2003 in paleontology

List of years in paleontology (table)
In science
2000
2001
2002
2003
2004
2005
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Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2003.

Plants

[edit]

Conifers

[edit]
Name Novelty Status Authors Age Unit Location Notes Images

Chamaecyparis eureka[2]

Sp nov

Valid

Kotyk, Basinger, & McIlver

Lutetian

Buchanan Lake Formation, Axel Heiberg Island

 Canada
 Nunavut

the oldest confirmed species of Chamaecyparis[2]

Tsuga swedaea[3]

Sp nov

Valid

LePage

Lutetian

Buchanan Lake Formation, Axel Heiberg Island

 Canada
 Nunavut

A hemlock species

Angiosperms

[edit]
Name Novelty Status Authors Age Unit Location Notes Images

Araripia florifera[4]

Gen et sp

Valid

Mohr & Eklund

lower Cretaceous

Crato Formation

 Brazil

Carpinus perryae[5]

Sp nov

Valid

Pigg, Manchester, & Wehr

Eocene
Ypresian

Okanagan Highlands
Klondike Mountain Formation

 USA
 Washington

A hornbeam

Carpinus parryae

Corylus johnsonii[5]

Sp nov

Valid

Pigg, Manchester, & Wehr

Eocene
Ypresian

Okanagan Highlands
Klondike Mountain Formation

 USA
 Washington

A hazelnut

Corylus johnsonii

Palaeocarpinus barksdaleae[5]

Sp nov

Valid

Pigg, Manchester, & Wehr

Eocene
Ypresian

Okanagan Highlands
Klondike Mountain Formation

 USA
 Washington

A betulaceous fruit

Palaeocarpinus barksdaleae

Palaeocarpinus dentatus[5]

Comb nov

Valid

(Penhallow) Pigg, Manchester, & Wehr

Eocene
Ypresian

Okanagan Highlands
"Stump lake", Coldwater Beds

 Canada
 British Columbia

A betulaceous fruit;
moved from Carpolithes dentatus (1890)

Palaeocarpinus stonebergae[5]

Sp nov

Valid

Pigg, Manchester, & Wehr

Eocene
Ypresian

Okanagan Highlands
Allenby Formation

 Canada
 British Columbia

A betulaceous fruit

Pseudosalix[6]

Gen et sp nov

Valid

Boucher, Manchester, & Judd

latest Ypresian

Green River Formation

 USA
 Utah

A willow relative

Gnetophytes

[edit]
Name Novelty Status Authors Age Unit Location Notes Images

Cratonia cotyledon[7]

Gen et sp nov

Valid

Rydin, Mohr, & Friis

lower Cretaceous

Crato Formation

 Brazil

A relative to Welwetschia

Fungi

[edit]
Name Novelty Status Authors Age Unit Location Notes Images

Aureofungus[8]

Get et sp nov

Valid

Hibbett et al

Burdigalian

Dominican amber.

 Dominican Republic

An agaricalean fungus
Type species A. yaniguaensis

Arthropods

[edit]

Arachnids

[edit]
Name Novelty Status Authors Age Unit Location Notes Images

Cornupalpatum[9]

Gen et sp nov

valid

Poinar & Brown

Albian-Cenomanian

Burmese amber

 Myanmar

An Ixodid hard tick

Cornupalpatum burmanicum

Insects

[edit]
Name Novelty Status Authors Age Unit Location Notes Images

Archimyrmex piatnitzkyi[10]

Comb nov.

vaild

(Viana & Haedo Rossi)

Middle Eocene

Ventana Formation

 Brazil

A myrmeciine ant.
moved from Ameghinoia piatnitzkyi (1957)

Archimyrmex piatnitzkyi

Archimyrmex smekali[10]

Comb nov.

valid

(Rossi de Garcia)

Middle Eocene

Ventana Formation

 Brazil

A myrmeciine ant.
moved from Polanskiella smekali (1983)

Brownimeciinae[11]

Subfam. Gen. et Sp. nov

Valid

Bolton

Turonian

New Jersey amber

 USA
 New Jersey

A stem group ant, type species B. clavata

Brownimecia clavata

Halictus? savenyei[12]

Sp. nov

valid

Engel & Archibald

Ypresian

Coldwater Beds

 Canada
 British Columbia

first apoid bee fossil described from Canada

Lasioglossum celinae[13]

Sp. nov

Valid

Nel & Petrulevicius

Rupelian

Dauphin

 France

A sweat bee

Lasioglossum celinae

Oligoapis[13]

Gen et sp nov

Jr synonym

Nel & Petrulevicius

Aquitanian

Bes-Konak

 Turkey

An apine bee,
type species O. beskonakensis
Moved to Bombus beskonakensis in 2019[14]

Bombus beskonakensis

Palaeopsychops dodgeorum[15]

Sp nov

valid

Makarkin & Archibald

Ypresian

Coldwater Beds

 Canada
 British Columbia

An ithionid giant lacewing

Paraelectrobombus[13]

Gen et sp nov

Jr synonym

Nel & Petrulevicius

Aquitanian

Bes-Konak

 Turkey

Described as an Electrobombini bee
type species P. patriciae
Moved to Bombus (Paraelectrobombus) patriciae in (2019)

Bombus (Paraelectrobombus) patriciae

Raphidia funerata[16]

sp nov

synonym

Engel

Priabonian

Florissant Formation

 USA
 Colorado

A snakefly.
moved to Florissantoraphidia funerata in 2014

Wesmaelius mathewesi[17]

Sp. nov

valid

Engel & Archibald

Ypresian

Coldwater beds, Princeton Group

 Canada
( British Columbia)

A Hemerobiinae lacewing

Xylocopa celineae[13]

Sp nov

valid

Nel & Petrulevicius

Chattian

Camoins-les-Bains

 France

A carpenter bee

Xylocopa celineae

Conodont paleozoology

[edit]

German paleontologist and stratigrapher Heinz Walter Kozur (1942-2013) described the conodont genus Carnepigondolella.

Vertebrate paleozoology

[edit]

Parareptiles

[edit]
Name Novelty Status Authors Age Unit Location Notes Images

Bunostegos

Valid

Sidor

 Niger

This was one of the largest herbivores of their time

Bunostegos

Non-avian dinosaurs

[edit]

Data courtesy of George Olshevsky's dinosaur genera list.[18]

Name Status Authors Location Notes Images
"Abdallahsaurus"[19]

Nomen nudum.

Nomen nudum; possibly a junior synonym of Giraffatitan or Brachiosaurus

"Blancocerosaurus"[20]

Nomen nudum.

Nomen nudum; possibly a junior synonym of Giraffatitan or Brachiosaurus

Amazonsaurus[21] Valid taxon
  • Carvalho
  • Avilla
  • Salgado
 Brazil
Antetonitrus[22] Valid taxon
  • Yates
  • Kitching
 South Africa
Aviatyrannis[23] Valid taxon
  • Rauhut
 Portugal
Bainoceratops[24] Valid taxon
  • Tereschenko
  • Alifanov
 China
Brohisaurus Valid taxon
  • Malkani
 Pakistan
Colepiocephale[25] Valid taxon
  • Sullivan
 Canada
"Domeykosaurus" Nomen nudum

Rubilar-Rogers and Vargas vide:

  • Anonymous on May 29, 2003 Reuters news reports.
 Chile
Dromaeosauroides[26] Valid taxon
  • Christiansen
  • Bonde
 Denmark
Dromaeosauroides
Equijubus[27] Valid taxon
  • You
  • Luo
  • Shubin
  • Witmer
  • Tang Z.
  • Tang F.
 China
Ferganasaurus[28] Valid taxon
  • Alifanov
  • Averianov
 Kyrgyzstan
Fukuisaurus[29] Valid taxon
  • Kobayashi
  • Azuma
 Japan
Gobititan[30] Valid taxon
  • You
  • Tang F.
  • Luo
 China
Hanssuesia[25] Valid taxon
  • Sullivan
 Canada ( Alberta)
Hanssuesia
Heyuannia[31] Valid taxon
 China
Heyuannia
Hongshanosaurus[32] Valid taxon
  • You
  • Xu X.
  • Wang X.
 China
Hongshanosaurus
Isisaurus[33] Valid taxon  India
Isisaurus
"Issasaurus"[19]

Nomen nudum; possibly a junior synonym of Dicraeosaurus

Lamaceratops[34] Valid taxon
  • Alifanov
 China
"Ligomasaurus"[19]

Nomen nudum; possibly a junior synonym of Giraffatitan or Brachiosaurus

Lusotitan[35] Valid taxon
  • Antunes
  • Mateus
 Portugal
Magnirostris[36] Valid taxon  China
Magnirostris
Mendozasaurus[37] Valid taxon
  • González-Riga (as González Riga)
 Argentina
"Mohammadisaurus"[19]

Nomen nudum.

Nomen nudum; possibly a junior synonym of Tornieria

"Mtapaiasaurus"[19]

Nomen nudum.

Nomen nudum; possibly a junior synonym of Giraffatitan or Brachiosaurus

"Mtotosaurus"[19]

Nomen nudum.

Nomen nudum; possibly a junior synonym of Dicraeosaurus

"Nteregosaurus"[19]

Nomen nudum.

Nomen nudum; possibly a junior synonym of Janenschia

"Nyororosaurus"[19]

Nomen nudum; possibly a junior synonym of Dicraeosaurus

Olorotitan[38] Valid taxon
  • Godefroit
  • Bolotsky
  • Alifanov
 Russia
Olorotitan
Olorotitan
Platyceratops[34] Valid taxon
  • Alifanov
 Mongolia
Rajasaurus[39] Valid taxon  India
Rajasaurus
Rinconsaurus[40] Valid taxon
  • Calvo
  • González-Riga (as González Riga)
 Argentina
Rinconsaurus
"Salimosaurus"[19]

Nomen nudum.

Nomen nudum; possibly a junior synonym of Giraffatitan or Brachiosaurus

"Selimanosaurus"[19]

Nomen nudum.

Nomen nudum; possibly a junior synonym of Dicraeosaurus

Serendipaceratops[41] Valid taxon
  • T. Rich
  • Vickers-Rich
 Australia
Shenzhousaurus[42] Valid taxon  China
Shuangmiaosaurus[43] Valid taxon
  • You
  • Ji Q.
  • Li J.
  • Li Y. X.
 China
Silesaurus[44] Valid taxon
  • Dzik
 Poland
Silesaurus
Sinornithomimus[45] Valid taxon
  • Kobayashi
 China
Sinornithomimus
Sphaerotholus[46] Possible junior synonym of Prenocephale
  • Williamson
  • Carr
"Wangonisaurus"[19]

Nomen nudum.

Nomen nudum; possibly a junior synonym of Giraffatitan or Brachiosaurus

Yixianosaurus[47] Valid taxon
  • Xu X.
  • Wang X.
 China
Zalmoxes[48] Valid taxon
  • Weishampel
  • Jianu
  • Csiki
  • Norman
 Romania
Zalmoxes
Zupaysaurus[49] Valid taxon

Arcucci & Coria

 Argentina
Zupaysaurus

Newly named birds

[edit]
Name Status Novelty Authors Age Unit Location Notes Images

Amanuensis pickfordi [50]

Gen. et sp. nov

valid

Cécile Mourer-Chauviré

Middle Miocene

MN 4

 Namibia

A Sagittariidae, relative of the secretarybird. The type species of Amanuensis Mourer-Chauviré, 2003.

Amitabha urbsinterdictensis [51]

Valid

Gen. nov. et Sp. nov.

Bonnie E. Gulas-Wroblewski

Anton F.-J. Wroblewski

Middle Eocene

Bridger Formation

 USA:

 Wyoming

The type species of Amitabha Gulas-Wroblewski & Wroblewski, 2003. Placed in crown Galliformes, but only compared with Phasianidae, not with other Eocene birds from North America.

Anhinga minuta [52]

Valid

Sp. nov.

Herculano M. F. de Alvarenga

Edson Guilherme

Late Miocene-Early Pliocene

Huayquerian-Montehermosian,

Solimõas Formation

 Brazil

An Anhingidae.

Coenocorypha miratropica [53]

Valid

Sp. nov.

Trevor H. Worthy

Holocene

Vitilevu

 Fiji

A Scolopacidae.

Diomedea tanakai [54]

Valid

Sp. nov.

Paul G. Davis

Late Early Miocene

Oi Formation

 Japan

A Diomedeidae.

Eoeurypyga olsoni [55]

Nomen Nudum

Thesis name.

Ilka Weidig

Early Eocene

Green River Formation

 USA:

 Wyoming

A Eurypygidae.

Falco umanskajae [56]

Valid

Sp. nov.

Denis V. Sobolev

Pliocene

MN 16

 Ukraine

A Falconidae.

Gallicolumba leonpascoi [57]

Valid

Sp. nov.

Trevor H. Worthy

Graham M. Wragg

Late Pleistocene-Holocene

Henderson Island

 Pitcairn

A Columbidae.

Hemignathus vorpalis [58]

Valid

Sp. nov.

Helen F. James

Storrs L. Olson

Subresent

Hawaii

 USA:

 Hawaii

A Fringillidae, Carduelinae, Drepanidini.

Macranhinga ranzii [52]

Valid

Sp. nov.

Herculano M. F. de Alvarenga

Edson Guilherme

Late Miocene-Early Pliocene

Huayquerian-Montehermosian

 Brazil

An Anhingidae.

Milvago carbo [59]

Valid

Sp. nov.

William Suárez Duque

Storrs L. Olson

Quaternary, possibly Late Pleistocene or Early Holocene

Las Breas de San Felipe

 Cuba

A Falconidae.

Morus peruvianus [60]

Valid

Sp. nov.

Marcelo Stucchi

Pliocene

Pisco Formation

 Peru

A Sulidae.

Namibiavis senutae[50]

Gen. et sp. nov

valid

Cécile Mourer-Chauviré

Middle Miocene

MN 4

 Namibia

Initially thought to be an Idiornithidae, Cariamiformes but in 2011 reinterpreted as an Opisthocomidae, a relative of the hoatzin.[61] The type species of Namibiavis Mourer-Chauviré, 2003.

Nycticorax olsoni [62]

Valid

Sp. nov.

W. R. P. Bourne

N. P. Ashmole

K. E. L. Simmons

Subrecent

Ascension Island

 Ascension Island

An Ardeidae.

Palaeogrus mainburgensis [63]

Valid

Sp. nov.

Ursula B. Göhlich

Early Middle Miocene (MN 5)

Upper Freshwater Molasse

 Germany

 Bavaria

A Gruidae.

Parargornis messelensis [64]

Valid

Gen. nov et Sp. nov.

Gerald Mayr

Early Middle Eocene

Messel pit,

MP 11

 Germany

 Hessen

An Apodiformes, Cypselavidae Mourer-Chauviré, 2006.

Shandongornis yinansis [65]

Valid

Gen. nov et Sp. nov.

Hou Lianhai

Pliocene

 China

A Phasianidae.

Spheniscus chilensis [66]

Valid

Sp. nov.

Steven D. Emslie

Carlos Guerra Correa

Pliocene

Mejillones Formation

 Chile

A Spheniscidae.

Spheniscus megaramphus [67]

Valid

Sp. nov.

Marcello Stucchi

Mario Urbina

Alfredo Giraldo

Late Miocene

Pisco Formation

 Peru

A Spheniscidae.

Sula magna [60]

Valid

Sp. nov.

Marcello Stucchi

Late Miocene-Early Pliocene

Pisco Formation

 Peru

A Sulidae.

Sula sulita [60]

Valid

Sp. nov.

Marcello Stucchi

Late Miocene-Early Pliocene

Pisco Formation

 Peru

A Sulidae.

Wyomingcypselus pohli [55]

Nomen Nudum

Thesis name.

Ilka Weidig

Early Eocene

Green River Formation

 USA:

 Wyoming

An Apodiformes.

Plesiosaurs

[edit]
Name Status Authors Notes

Terminonatator

Valid

Sato

Thililua

Valid

Bardet Suberbiola Jalil

Pterosaurs

[edit]
Name Status Authors Notes

Beipiaopterus

Valid

Lü J.-C.

Chaoyangopterus

Valid

Wang Zhou

Harpactognathus

Valid

Carpenter Unwin et al.

Jidapterus

Valid

Dong, Sun, & Wu,

Liaoningopterus

Valid

Wang X.-L. and Zhou Z.-H

Liaoningopteryx

Valid

Wang X. Zhao

Ludodactylus

Valid

Frey, E., Martill Buchy, M

Nurhachius

Valid

Wang Kellner et al.

Phosphatodraco

Valid

Pereda-Suberbiola N. Bardet et al.

Sinopterus

Valid

Wang, X. Zhou, Z

Synapsids

[edit]

Non-mammalian

[edit]
Name Status Authors Discovery year Age Unit Location Notes Images

Archaeovenator[68]

Valid

  • Reisz
  • Dilkes

Upper Carboniferous
(Upper Pennsylvanian)

Hamilton Quarry

 USA

a pelycosaur

Brasilitherium[69]

Valid

  • Bonaparte
  • Martinelli
  • Schultz
  • Rupert

 Brazil

Brasilodon[69]

Valid

  • Bonaparte
  • Martinelli
  • Schultz
  • Rupert

 Brazil

Bullacephalus[70]

Valid

  • Rubidge
  • Kitching

 South Africa

Colobodectes[71]

Valid

  • Modesto
  • Rubidge
  • Visser
  • Welman

Upper/Middle Permian
(Capitanian)

Abrahamskraal formation

 South Africa

an anomodont

Lanthanostegus[72]

Valid

  • Modesto
  • Rubidge
  • Welman

Middle/Upper Permian
(Capitanian)

 South Africa

an anomodont; replacement name for Lanthanocephalus Modesto, Rubidge & Welman, 2002, preoccupied by the cnidarian genus Lanthanocephalus Williams & Starmer, 2000[73]

Santacruzodon[74]

Valid

  • Abdala
  • Ribeiro

 Brazil

Xiyukannemeyeria[75]

Valid

  • Liu
  • Li

Middle Triassic
(Anisian)

 China

an anomodont; new genus for "Parakannemeyeria" brevirostris

Mammals

[edit]
Name Novelty Status Authors Age Unit Location Notes Images

Caracal depereti[76]

Sp nov

Disputed

Morales et al.

Pliocene

 Spain

A Caracal. Boscaini et al. (2016) considered this species to be a junior synonym of Lynx issiodorensis.[77]

Guangxicyon[78] Gen. et sp. nov. Valid Zhai et al. Eocene Nadu Formation  China An amphicyonid.

Iridodon[79]

Gen et sp nov

Valid

Morlo & Gunnell

Bridgerian

Wasatch Formation

 United States

A limnocyonid hyaenodont.The type species is I. datzae.

Kogolepithecus[80]

Gen et sp nov

Valid

Pickford et al.

Miocene

 Uganda

A catarrhine of uncertain affinity. The type species is K. morotoensis.

Yuesthonyx[81] Gen. et sp. nov. Valid Tong, Wang & Fu Paleocene Dazhang Formation  China A tillodont. The type species is Y. tingae.

References

[edit]
  1. ^ Gini-Newman, Garfield; Graham, Elizabeth (2001). Echoes from the past: world history to the 16th century. Toronto: McGraw-Hill Ryerson Ltd. ISBN 9780070887398. OCLC 46769716.
  2. ^ a b Kotyk, M.E.A.; Basinger, J.F.; McIlver, E.E. (2003). "Early Tertiary Chamaecyparis Spach from Axel Heiberg Island, Canadian High Arctic". Canadian Journal of Botany. 81 (2): 113–130. doi:10.1139/B03-007.
  3. ^ Lepage, B. A. (2003). "A new species of Tsuga (Pinaceae) from the middle Eocene of Axel Heiberg Island, Canada, and an assessment of the evolution and biogeographical history of the genus". Botanical Journal of the Linnean Society. 141 (3): 257–296. doi:10.1046/j.1095-8339.2003.00131.x.
  4. ^ Mohr, B.A.R.; Eklund, H. (2003). "Araripia florifera, a magnoliid angiosperm from the Lower Cretaceous Crato Formation (Brazil)". Review of Palaeobotany and Palynology. 126 (3–4): 279–292. Bibcode:2003RPaPa.126..279M. doi:10.1016/S0034-6667(03)00092-7.
  5. ^ a b c d e Pigg, K.B.; Manchester S.R.; Wehr W.C. (2003). "Corylus, Carpinus, and Palaeocarpinus (Betulaceae) from the Middle Eocene Klondike Mountain and Allenby Formations of Northwestern North America". International Journal of Plant Sciences. 164 (5): 807–822. doi:10.1086/376816. S2CID 19802370.
  6. ^ Boucher, L. D.; Manchester, S.; Judd, W. (2003). "An extinct genus of Salicaceae based on twigs with attached flowers, fruits, and foliage from the Eocene Green River Formation of Utah and Colorado, USA". American Journal of Botany. 90 (9): 1389–99. doi:10.3732/ajb.90.9.1389. PMID 21659238.
  7. ^ Rydin, C.; Mohr, B.; Friis, E.M. (2003). "Cratonia cotyledon gen. et sp. nov.: a unique Cretaceous seedling related to Welwetschia". Proceedings of the Royal Society of London B. 270 (Suppl 1): S29–S32. doi:10.1098/rsbl.2003.0044. PMC 1698015. PMID 12952628.
  8. ^ hibbet, D.S.; et al. (2003). "Another Fossil Agaric from Dominican Amber". Mycologia. 95 (4): 685–687. doi:10.2307/3761943. JSTOR 3761943. PMID 21148976.
  9. ^ Poinar, G.; Brown, A. E. (2003). "A new genus of hard ticks in Cretaceous Burmese amber (Acari: Ixodida: Ixodidae)". Systematic Parasitology. 54 (3): 199–205. doi:10.1023/A:1022689325158. PMID 12652071. S2CID 11527194.
  10. ^ a b Dlussky, G.M.; Perfilieva, K.S. (2003). "Paleogene ants of the genus Archimyrmex Cockerell, 1923 (Hymenoptera, Formicidae, Myrmeciinae)" (PDF). Paleontological Journal. 37 (1): 39–47.
  11. ^ Bolton, B. (2003), "Synopsis and classification of Formicidae.", Memoirs of the American Entomological Institute, 71: 1–370
  12. ^ Engel, Michael S.; Archibald, S. Bruce (2003). "An Early Eocene bee (Hymenoptera: Halictidae) from Quilchena, British Columbia". The Canadian Entomologist. 135: 63–69. doi:10.4039/n02-030. hdl:1808/16473. S2CID 54053341.
  13. ^ a b c d Nel, A; Petrulevicius, J. F. (2003). "New Palaeogene bees from Europe and Asia". Alcheringa. 27 (4): 277–293. Bibcode:2003Alch...27..277N. doi:10.1080/03115510308619108. S2CID 84242180.
  14. ^ Dehon, Manuel; Engel, Michael S.; Gérard, Maxence; Aytekin, A. Murat; Ghisbain, Guillaume; Williams, Paul H.; Rasmont, Pierre; Michez, Denis (2019-11-21). "Morphometric analysis of fossil bumble bees (Hymenoptera, Apidae, Bombini) reveals their taxonomic affinities". ZooKeys (891). Pensoft Publishers: 71–118. Bibcode:2019ZooK..891...71D. doi:10.3897/zookeys.891.36027. ISSN 1313-2970. PMC 6882928. PMID 31802973.
  15. ^ Makarkin, V. N.; Archibald, S. B. (2003). "Family affinity of the genus Palaeopsychops Andersen with description of a new species from the Early Eocene of British Columbia, Canada (Neuroptera: Polystoechotidae)". Annals of the Entomological Society of America. 96 (3): 171–180. doi:10.1603/0013-8746(2003)096[0171:FAOTGP]2.0.CO;2. S2CID 84362010.
  16. ^ Engel, M.S. (2003). "A New Eocene-Oligocene Snakefly from Florissant, Colorado (Raphidioptera: Raphidiidae)". Transactions of the Kansas Academy of Science. 106 (3 & 4): 124–128. doi:10.1660/0022-8443(2003)106[0124:anesff]2.0.co;2. JSTOR 3628392. S2CID 85677413.
  17. ^ Makarkin, VN; Archibald, SB; Oswald, JD (2003). "New Early Eocene brown lacewings (Neuroptera: Hemerobiidae) from western North America". The Canadian Entomologist. 135 (5): 637–653. CiteSeerX 10.1.1.489.5852. doi:10.4039/n02-122. S2CID 53479449.
  18. ^ Olshevsky, George. "Dinogeorge's Dinosaur Genera List". Archived from the original on 2011-07-15. Retrieved 2008-08-07.
  19. ^ a b c d e f g h i j k Maier, G. (2003). African Dinosaurs Unearthed: The Tendaguru Expeditions. Life of the Past. Indiana University Press. ISBN 978-0-253-00054-5.
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