2013 in arthropod paleontology

List of years in arthropod paleontology
In paleontology
2010
2011
2012
2013
2014
2015
2016
In science
2010
2011
2012
2013
2014
2015
2016
In reptile paleontology
2010
2011
2012
2013
2014
2015
2016
In paleobotany
2010
2011
2012
2013
2014
2015
2016
In paleoentomology
2010
2011
2012
2013
2014
2015
2016
In paleomalacology
2010
2011
2012
2013
2014
2015
2016
In archosaur paleontology
2010
2011
2012
2013
2014
2015
2016
In mammal paleontology
2010
2011
2012
2013
2014
2015
2016
In paleoichthyology
2010
2011
2012
2013
2014
2015
2016

This list of fossil arthropods described in 2013 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods of every kind that have been described during the year 2013. The list only includes taxa at the level of genus or species.

Arachnids

[edit]
Name Novelty Status Authors Age Unit Location Notes Images

Amiracarus pliocennatus[1]

Sp. nov

Valid

Miko et al.

Pliocene to Pleistocene

 Slovenia

An oribatid mite.

Chaerilobuthus longiaculeus[2]

Sp. nov

Valid

Lourenço

Cenomanian

Burmese amber

 Myanmar

A scorpion, a species of Chaerilobuthus.

Friularachne[3]

Gen. et sp. nov

Valid

Dalla Vecchia & Selden

Late Triassic (Norian)

Dolomia di Forni Formation

 Italy

A member of Mygalomorphae, possibly an atypoid. The type species is Friularachne rigoi.

Gondwanascorpio[4]

Gen. et sp. nov

Valid

Gess

Devonian (Famennian)

Witpoort Formation

 South Africa

A scorpion of uncertain phylogenetic placement. The type species is Gondwanascorpio emzantsiensis.

Labidostoma (Pseudocornutella) electri[5]

Subgen. et sp. nov

Valid

Sidorchuk & Bertrand

Eocene

Europe

A labidostomatid mite found in Baltic amber, a species of Labidostoma.

Lagonomegops? cor[6]

Sp. nov

Valid[7]

Pérez-de la Fuente, Saupe & Selden

Early Cretaceous (Albian)

 Spain

A lagonomegopid found in Albian Spanish amber, possibly a species of Lagonomegops.

Mongolarachne[8]

Gen. et comb. nov

Valid

Selden, Shih & Ren

Middle or Late Jurassic

 China

A member of Araneomorphae related to the group Orbiculariae; a new genus for "Nephila" jurassica Selden, Shih & Ren (2011).

Permotarbus[9]

Gen. et sp. nov

Valid

Dunlop & Rößler

Early Permian

 Germany

A member of Trigonotarbida. The type species is Permotarbus schuberti.

Sellnickiella balticae[5]

Sp. nov

Valid

Sidorchuk & Bertrand

Eocene

Europe

A labidostomatid mite found in Baltic amber, a species of Sellnickiella.

Soplaogonomegops[6]

Gen. et sp. nov

Valid[7]

Pérez-de la Fuente, Saupe & Selden

Early Cretaceous (Albian)

 Spain

A lagonomegopid found in Albian Spanish amber. The type species is Soplaogonomegops unzuei.

Spinomegops[6]

Gen. et 2 sp. nov

Valid[7]

Pérez-de la Fuente, Saupe & Selden

Early Cretaceous (Albian)

 Spain

A lagonomegopid found in Albian Spanish amber. Genus contains two species: S. arcanus and S. aragonensis.

Talbragaraneus[10]

Gen. et sp. nov

Valid

Selden & Beattie

Jurassic

Talbragar Fossil Fish Bed

 Australia

A possible member of Uloboridae. The type species is Talbragaraneus jurassicus.

Tityus azari[11]

Sp. nov.

Disputed

Lourenço

Late Eocene or early Oligocene

 Dominican Republic

A buthid scorpion found in Dominican amber, a species of Tityus. Teruel (2017) considered this species to be a junior synonym of Tityus geratus Santiago-Blay & Poinar (1988),[12] while Lourenço (2017) considered it to be a distinct species.[13]

Tynecotarbus[14]

Gen. et sp. nov

Valid

Hradská & Dunlop

Carboniferous (late Pennsylvanian)

 Czech Republic

A member of Trigonotarbida. The type species is Tynecotarbus tichaveki.

Crustaceans

[edit]
Name Novelty Status Authors Age Unit Location Notes Images

Acareprosopon[15]

Gen. et comb. nov

Valid

Klompmaker

Early Cretaceous (late Albian)

Eguino Formation

 Spain

A crab belonging to the family Prosopidae. The type species is "Pithonoton" bouvieri Van Straelen (1944).

Acratia? hamadai[16]

Sp. nov

Valid

Tanaka, Ono & Maeda in Tanaka et al.

Middle Permian

 Japan

An ostracod, possibly a species of Acratia.

Acratia? jeanvannieri[17]

Sp. nov

Valid

Forel in Forel et al.

Late Permian to Early Triassic

 Hungary

An acratiid ostracod, a possible species of Acratia.

Acratia nagyvisnyoensis[17]

Sp. nov

Valid

Forel in Forel et al.

Late Permian (Changhsingian)

Nagyvisnyó Limestone Formation

 Hungary

An acratiid ostracod, a species of Acratia.

Acratia? okumurai[16]

Sp. nov

Valid

Tanaka in Tanaka et al.

Middle Permian

 Japan

An ostracod, possibly a species of Acratia.

Aeger luxii[18]

Sp. nov

Valid

Huang et al.

Middle Triassic

Guanling Formation

 China

A penaeoid, a species of Aeger.

Aegyptoleberis[19]

Gen. et comb. nov

Valid

Boukhary, Bassiouni & Sharabi in Boukhary et al.

Paleocene

 Egypt

An ostracod; a new genus for "Cythereis" coronata Esker (1968).

Albaidaplax[20]

Gen. et sp. nov

Valid

Garassino, Pasini & Castro

Pliocene to early Pleistocene

 Italy
 Spain

A goneplacid crab. The type species is Albaidaplax ispalensis. Announced in 2013; validated in 2023.[21]

Albenizus[15]

Gen. et sp. nov

Valid

Klompmaker

Early Cretaceous (late Albian)

Eguino Formation

 Spain

A crab belonging to the family Torynommidae. The type species is Albenizus minutus.

Ambulocapsa[22]

Gen. et 4 sp. nov

Valid

Robins, Feldmann & Schweitzer

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A munidopsid galatheoid. Genus contains four species: Ambulocapsa altilis, Ambulocapsa bachmayeri, Ambulocapsa novacula and Ambulocapsa sentosa.

Antrimpos germanicus[23]

Sp. nov

Valid

Brandt & Schulz

Middle Triassic

Trochitenkalk Formation

 Germany

A member of the family Penaeidae. Originally described as a species of Antrimpos, but subsequently transferred to the genus Gladiocaris.[24]

Amphicytherura fragilis[25]

Sp. nov

Valid

Nicolaidis & Piovesan in Piovesan et al.

Early Cretaceous (Albian)

São Mateus Formation

 Brazil

An ostracod belonging to the family Cytheridae. Originally described as a species of Amphicytherura;[25] Antonietto et al. (2016) transferred the species to the genus Aracajuia.[26]

Angarestia[27]

Gen. et comb. et 3 sp. nov

Valid

Charbonnier et al.

Jurassic and Cretaceous

 Antarctica
 Australia
 Italy
 Lebanon

A member of the family Glypheidae. A new genus for "Glyphea" australensis Feldmann, Tshudy & Thomson (1993); genus also includes "Glyphea" tricarinata Garassino (1996), "Glyphea" foresti Feldmann & de Saint Laurent (2002), and "Glyphea" damesi Garassino (2001), as well as new species A. viallei, A. anatoliensis and A. mugudaensis.

Ankylokypha[22]

Gen. et sp. nov

Valid

Robins, Feldmann & Schweitzer

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A munidopsid galatheoid. The type species is Ankylokypha parabola.

Archaeoportunus[28]

Gen. et sp. nov

Valid

Artal, Ossó & Domínguez

Early Eocene (Ypresian)

Roda Formation

 Spain

A crab, a member of Portunoidea. The type species is Archaeoportunus isabenensis.

Archaeotetra lessinea[29]

Sp. nov

Valid

De Angeli & Ceccon

Eocene (Ypresian)

 Italy

A trapeziid trapezioid, a species of Archaeotetra.

Arcoscalpellum concavitectum[30]

Sp. nov

Valid

Carriol

Pliocene

 France

A barnacle, a species of Arcoscalpellum.

Astiplax[31]

Gen. et sp. nov

Valid

Garassino & Pasini

Late Pliocene

 Italy

A goneplacid crab. The type species is Astiplax aspera.

Audogaster[27]

Gen. et comb. et sp. nov

Valid

Charbonnier et al.

Triassic

 Germany

A member of Glypheoidea belonging to the family Litogastridae. Genus includes "Pseudopemphix" spinosus Assmann (1927), as well as new species A. assmanni.

Aulavescus[22]

Gen. et 2 sp. nov

Valid

Robins, Feldmann & Schweitzer

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A munidopsid galatheoid. Genus contains two species: Aulavescus exutus and Aulavescus tectus.

Aurikirkbya kinshozanensis[16]

Sp. nov

Valid

Tanaka in Tanaka et al.

Middle Permian

 Japan

An ostracod, a species of Aurikirkbya.

Aysegulina mbassisensis[32]

Sp. nov

Valid

Sarr

Paleocene (Thanetian)

 Senegal

An ostracod belonging to the family Trachyleberididae.

Aysegulina papillata[32]

Sp. nov

Valid

Sarr

Paleocene-Eocene (Thanetian-Lutetian)

 Guinea-Bissau
 Senegal

An ostracod belonging to the family Trachyleberididae.

Bachmayerus[33]

Gen. et sp. nov

Valid

Fraaije et al.

Late Jurassic to Early Cretaceous (Tithonian to early Berriasian)

Štramberk Limestone

 Czech Republic

A hermit crab, a member of Diogenidae. The type species is Bachmayerus cavus.

Bairdia akasakaensis[16]

Sp. nov

Valid

Tanaka in Tanaka et al.

Middle Permian

 Japan

An ostracod, a species of Bairdia.

Bairdia anisongae[17]

Sp. nov

Valid

Forel in Forel et al.

Late Permian to Early Triassic

 Hungary

A bairdiid ostracod, a species of Bairdia.

Bairdia baudini[17]

Nom. nov

Valid

Crasquin in Forel et al.

Late Permian to Early Triassic

 China
 Hungary
 Turkey

A replacement name for the bairdiid ostracod species Bairdia subsymmetrica (Shi in Shi and Chen, 1987).

Bairdia davehornei[17]

Sp. nov

Valid

Forel in Forel et al.

Late Permian to Early Triassic

 Hungary

A bairdiid ostracod, a species of Bairdia.

Bairdia nishiwakii[16]

Sp. nov

Valid

Tanaka & Nishimura in Tanaka et al.

Middle Permian

 Japan

An ostracod, a species of Bairdia.

Bairdia oogakiensis[16]

Sp. nov

Valid

Tanaka in Tanaka et al.

Middle Permian

 Japan

An ostracod, a species of Bairdia.

Bairdiacypris chaabetensis[34]

Sp. nov

Valid

Andreu et al.

Late Cretaceous

 Morocco

An ostracod.

Bairdiacypris? hayasakai[16]

Sp. nov

Valid

Tanaka in Tanaka et al.

Middle Permian

 Japan

An ostracod, possibly a species of Bairdiacypris.

Bathycalliax mediterranea[35]

Sp. nov

Valid

Baldanza et al.

Early Pleistocene

 Italy

A member of Callianassidae, a species of Bathycalliax.

Bittnerilia pentagonalis[36]

Sp. nov

Valid

Beschin, De Angeli & Zarantonello

Eocene

 Italy

A crab belonging to the family Parthenopidae.

Bonizzatoides[37]

Gen. et sp. nov

Valid

Beschin, Busulini & Tessier

Eocene

 Italy

A raninid crab. The type species is Bonizzatoides tuberculatus.

Boschettocheles[36]

Gen. et sp. nov

Valid

Beschin, De Angeli & Zarantonello

Eocene

 Italy

Genus includes new species B. magnum.

Brachycythere multidifferentis[25]

Sp. nov

Valid

Nicolaidis & Piovesan in Piovesan et al.

Late Cretaceous (Cenomanian-Turonian)

Florianópolis Formation
Itajaí-Açu Formation
Itanhaém Formation

 Brazil

An ostracod belonging to the family Brachycytheridae. Originally described as a species of Brachycythere; transferred to the genus Sapucariella by Puckett, Andreu & Colin (2016).[38]

Brasacypris subovatum[39]

Sp. nov

Valid

Do Carmo et al.

Early Cretaceous (Aptian)

Alagamar Formation

 Brazil

An ostracod, a species of Brasacypris.

Bulbileaia[40]

Gen. et sp. nov

Valid

Niu in Niu et al.

Late Permian

Taohaiyingzi Formation

 China

A clam shrimp. The type species is Bulbileaia orientalis.

Bullariscus[22]

Gen. et 4 sp. nov

Valid

Robins, Feldmann & Schweitzer

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A munidopsid galatheoid. Genus contains four species: Bullariscus arcuotorus, B. gibbernodus, B. patruliusi and B. triquetrus.

Bythocypris amelkisensis[34]

Sp. nov

Valid

Andreu et al.

Late Cretaceous

 Algeria[41]
 Morocco

An ostracod.

Calappilia minuta[37]

Sp. nov

Valid

Beschin, Busulini & Tessier

Eocene

 Italy

A calappid crab, a species of Calappilia.

Callianassa (s.l.) ocozocoautlaensis[42]

Sp. nov

Valid

Hyžný, Vega & Coutiño

Late Cretaceous (early Maastrichtian)

Ocozocoautla Formation

 Mexico

A member of Callianassidae, a species of Callianassa (sensu lato). Announced in 2013; validated in 2023.[43]

Callicythere? balvanyseptentrioensis[17]

Sp. nov

Valid

Forel in Forel et al.

Late Permian to Early Triassic

 Hungary

A bythocytherid ostracod, a possible species of Callicythere.

Candonocypris fimibolus[44]

Sp. nov

Valid[45]

Matzke-Karasz et al.

Early Miocene (23–16 Ma)

Riversleigh

 Australia

An ostracod, a species of Candonocypris.

Cavellina hashintotoi[16]

Sp. nov

Valid

Tanaka & Maeda in Tanaka et al.

Middle Permian

 Japan

An ostracod, a species of Cavellina.

Cenomanocarcinus nammourensis[46]

Sp. nov

Valid

Beschin & De Angeli

Late Cretaceous (Cenomanian)

 Lebanon

A raninoid crab, a species of Cenomanocarcinus.

Cenomanocarcinus robertsi[47]

Sp. nov

Valid

Feldmann et al.

Late Cretaceous (Campanian)

 United States

A raninoid crab, a species of Cenomanocarcinus.

Charybdis (Charybdis) kinugawai[48]

Sp. nov

Valid

Karasawa in Karasawa & Kinugawa

Late Pliocene

Ananai Formation

 Japan

A portunid crab, a species of Charybdis.

Chondromaia[47]

Gen. et sp. nov

Valid

Feldmann et al.

Late Cretaceous (early Maastrichtian)

Mount Laurel Formation

 United States

A majid crab. The type species is Chondromaia antiqua.

Cirolana feldmanni[49]

Sp. nov

Valid

Hyžný, Bruce & Schlögl

Early Miocene (late Karpatian, latest Burdigalian in the Mediterranean scale)

Lakšárska Nová Ves Formation

 Slovakia

A cirolanid isopod, a species of Cirolana.

Codoisopus[50]

Gen. et sp. nov

Valid

Lindoso, Carvalho & Mendes

Early Cretaceous (Aptian)

Codó Formation

 Brazil

An archaeoniscid isopod. The type species is Codoisopus brejensis.

Costa ? deklaszi[32]

Sp. nov

Valid

Sarr

Paleocene (Selandian-Thanetian)

 Senegal

An ostracod belonging to the family Trachyleberididae.

Cracensigillatus[22]

Gen. et 2 sp. et comb. nov

Valid

Robins, Feldmann & Schweitzer

Late Jurassic (Kimmeridgian[51] and Tithonian)

Ernstbrunn Limestone

 Austria
 Romania[51]

A munidopsid galatheoid. Genus contains two new species: Cracensigillatus gracilirostrus and Cracensigillatus prolatus, as well as the species Cracensigillatus acutirostrus (Moericke, 1889).

Cretamaja[15]

Gen. et sp. nov

Valid

Klompmaker

Early Cretaceous (late Albian)

Eguino Formation

 Spain

A crab belonging to the group Majoidea and the family Priscinachidae. The type species is Cretamaja granulata.

Cryptopagurus[52]

Gen. et sp. nov

Valid

Schweigert et al.

Early Jurassic (late Pliensbachian)

 Germany

A hermit crab. The type species is Cryptopagurus svenhofmanni.

Ctenocheles fabianii[37]

Sp. nov

Valid

Beschin, Busulini & Tessier

Eocene

 Italy

A member of Axiidea, a species of Ctenocheles.

Culmenformosa[22]

Gen. et 3 sp. nov

Valid

Robins, Feldmann & Schweitzer

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A munidopsid galatheoid. Genus contains three species: Culmenformosa glaessneri, Culmenformosa nephelepera and Culmenformosa triastrixosa.

Cycleryon bourseaui[53]

Sp. nov

Valid[54]

Audo et al.

Late Jurassic

 France

An eryonid, a species of Cycleryon.

Cytherella tazzouguertensis[34]

Sp. nov

Valid

Andreu et al.

Late Cretaceous

 Algeria[41]
 Morocco

An ostracod.

Cytherellina? magyarorszagensis[17]

Sp. nov

Valid

Forel in Forel et al.

Early Triassic (Griesbachian)

Gerennavár Limestone Formation

 Hungary

A bairdiocypridid ostracod, a possible species of Cytherellina.

Cytherelloidea saloumensis[32]

Sp. nov

Valid

Sarr

Paleocene-Eocene (Thanetian-Ypresian)

 Senegal

An ostracod belonging to the family Cytherellidae.

Cytheropteron dakhlaensis[19]

Sp. nov

Valid

Boukhary & Bassiouni in Boukhary et al.

Paleocene

 Egypt

An ostracod.

Cytheropteron mbeyensis[32]

Sp. nov

Valid

Sarr

Paleocene (Selandian-Thanetian)

 Senegal

An ostracod belonging to the family Cytheruridae.

Dahomeya reticulata[32]

Sp. nov

Valid

Sarr

Paleocene (Thanetian)

 Senegal

An ostracod belonging to the family Brachycytheridae.

Demania ahramensis[55]

Sp. nov

Valid

Yazdi et al.

Middle Miocene

Mishan Formation

 Iran

A xanthid crab, a species of Demania.

Dinglecythere[56]

Gen. et comb. nov

Valid

Antonietto et al.

Late Cretaceous (Santonian) to Eocene (Priabonian)

 Australia
 Jamaica
 South Africa
 United States

An ostracod belonging to the group Podocopida and the family Cytheridae. The type species is "Amphicytherura" tumida Dingle (1969); genus also includes "Amphicytherura" iniqua Holden (1964), "Amphicytherura" zululandensis Dingle (1980), "Amphicytherura" armatus Dingle (1981), "Amphicytherura" dinglei McKenzie et al. (1993) and "Amphicytherura" occulta Puckett & Colin in Puckett, Colin & Mitchell (2012).

Diogenes lessineus[36]

Sp. nov

Valid

Beschin, De Angeli & Zarantonello

Eocene

 Italy

A crab belonging to the family Diogenidae.

Discoidella niculaei[57]

Sp. nov

Valid

Sebe in Sebe, Crasquin & Grădinaru

Middle Triassic (Anisian)

 Romania

An ostracod belonging to the family Polycopidae.

Distefania lauginigeri[47]

Sp. nov

Valid

Feldmann et al.

Late Cretaceous (Campanian)

 United States

A goniodromitid homolodromioid crab, a species of Distefania.

Enoploclytia tepeyacensis[58]

Sp. nov

Valid

Vega, Garassino & Zapata Jaime

Late Cretaceous (Campanian)

 Mexico

An erymid, a species of Enoploclytia. Announced in 2013; validated in 2023.[59]

Eocytheropteron yoroensis[32]

Sp. nov

Valid

Sarr

Paleocene (Thanetian)

 Senegal

An ostracod belonging to the family Cytheruridae.

Eosestheria yaozhanensis[60]

Sp. nov

Valid

Wang

Late Jurassic

Dabeigou Formation

 China

A clam shrimp, a species of Eosestheria.

Etyus makrochele[61]

Sp. nov

Valid

Feldmann, Schweitzer & Lehmann

Early Cretaceous (late Albian)

 Germany

An etyiid crab, a species of Etyus.

Eumiraculum desmaresae[17]

Sp. nov

Valid

Forel in Forel et al.

Late Permian (Changhsingian)

Nagyvisnyó Limestone Formation

 Hungary

An ostracod of uncertain affinities, a species of Cytherellina.

Eurotetralia[29]

Gen. et comb. nov

Valid

De Angeli & Ceccon

Eocene

 Hungary
 Italy

A tetraliid trapezioid; a new genus for "Trapezia" loerenthey Müller (1975).

Fossocytheridea ballentae[25]

Sp. nov

Valid

Piovesan & Nicolaidis in Piovesan et al.

Cretaceous (Albian-Santonian)

Florianópólis Formation
Itajaí-Açú Formation

 Brazil

An ostracod belonging to the family Cytherideidae.

Fossocytheridea elegans[25]

Sp. nov

Valid

Piovesan & Nicolaidis in Piovesan et al.

Early Cretaceous (Albian)

Florianópólis Formation

 Brazil

An ostracod belonging to the family Cytherideidae.

Gastrodorus kotoucensis[33]

Sp. nov

Valid

Fraaije et al.

Late Jurassic to Early Cretaceous (Tithonian to early Berriasian)

Štramberk Limestone

 Czech Republic

A hermit crab, a member of Gastrodoridae; a species of Gastrodorus.

Gastrosacus aequabus[22]

Sp. nov

Valid

Robins, Feldmann & Schweitzer

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A munidopsid galatheoid, a species of Gastrosacus.

Gastrosacus levocardiacus[22]

Sp. nov

Valid

Robins, Feldmann & Schweitzer

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A munidopsid galatheoid, a species of Gastrosacus.

Gastrosacus limacurvus[22]

Sp. nov

Valid

Robins, Feldmann & Schweitzer

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A munidopsid galatheoid, a species of Gastrosacus.

Gastrosacus pisinnus[22]

Sp. nov

Valid

Robins, Feldmann & Schweitzer

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A munidopsid galatheoid, a species of Gastrosacus.

Gastrosacus torosus[22]

Sp. nov

Valid

Robins, Feldmann & Schweitzer

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A munidopsid galatheoid, a species of Gastrosacus.

Gaudipluma[62]

Gen. et sp. nov

Valid

Artal et al.

Eocene

 Spain

A retroplumid crab. The type species is Gaudipluma bacamortensis.

Gifuaparchites[16]

Gen. et sp. nov

Valid

Tanaka & Maeda in Tanaka et al.

Middle Permian

 Japan

An ostracod. The type species is Gifuaparchites takagii.

Gigacerina[27]

Gen. et comb. nov

Valid

Charbonnier et al.

Late Jurassic

 France

A member of the family Glypheidae. Genus includes "Glyphea" saemanni Oppel (1861).

Gigacerina saemanni

Glyphea macromuscula[63]

Sp. nov

Valid

Feldmann & Schweitzer

Early Jurassic

 United Kingdom

A glypheoid, a species of Glyphea.

Glypheopsis trouvillensis[27]

Sp. nov

Valid

Charbonnier et al.

Late Jurassic

 France

A member of the family Glypheidae.

Glypheopsis voultensis[27]

Sp. nov

Valid

Charbonnier et al.

Middle Jurassic

La Voulte-sur-Rhône

 France

A member of the family Glypheidae.

Glypturus persicus[64]

Sp. nov

Valid

Hyžný et al.

Middle-Late Miocene

 Iran

A callianassid, a species of Glypturus.

Harbinia crepata[39]

Sp. nov

Valid

Do Carmo et al.

Early Cretaceous (Aptian)

Alagamar Formation

 Brazil

An ostracod, a species of Harbinia.

Harbinia dimorphica[39]

Sp. nov

Valid

Do Carmo et al.

Early Cretaceous (Aptian)

Alagamar Formation

 Brazil

An ostracod, a species of Harbinia.

Hardapestheria[65]

Gen. et sp. nov

Valid[66]

Stigall et al.

Early Jurassic

Kalkrand Formation

 Namibia

An eosestheriid clam shrimp. The type species is Hardapestheria maxwelli.

Hazelina ? foundiougnensis[32]

Sp. nov

Valid

Sarr

Paleocene (Thanetian)

 Senegal

An ostracod belonging to the family Trachyleberididae.

Hemicycloleaia[40]

Gen. et sp. nov

Valid

Niu in Niu et al.

Late Permian

Taohaiyingzi Formation

 China

A clam shrimp. The type species is Hemicycloleaia curtus.

Hemiparacytheridea sagittaemucronata[34]

Sp. nov

Valid

Andreu et al.

Late Cretaceous

 Morocco

An ostracod.

Hermanites ? pustulosa[32]

Sp. nov

Valid

Sarr

Eocene

 Mauritania
 Senegal

An ostracod belonging to the family Trachyleberididae.

Heterocypris collaris[44]

Sp. nov

Valid[45]

Matzke-Karasz et al.

Early Miocene (23–16 Ma)

Riversleigh

 Australia

An ostracod, a species of Heterocypris.

Hollinella fengqinglaii[17]

Sp. nov

Valid

Crasquin in Forel et al.

Late Permian (Changhsingian)

Nagyvisnyó Limestone Formation

 Hungary

A hollinellid ostracod, a species of Hollinella.

Housacheles[33]

Gen. et sp. nov

Valid

Fraaije et al.

Late Jurassic to Early Cretaceous (Tithonian to early Berriasian)

Štramberk Limestone

 Czech Republic

A hermit crab, a member of Parapylochelidae. The type species is Housacheles timidus.

Hungarella gerennavarensis[17]

Sp. nov

Valid

Crasquin in Forel et al.

Early Triassic (Griesbachian)

Gerennavár Limestone Formation

 Hungary

A healdiid ostracod, a species of Hungarella.

Joeranina colombiana[67]

Sp. nov

Valid

Bermúdez et al.

Early Cretaceous (middle Albian)

 Colombia

A palaeocorystid decapod, a species of Joeranina.

Joeranina goshourajimensis[68]

Sp. nov

Valid

Karasawa & Komatsu

Early Cretaceous (late Albian)

Enokuchi Formation

 Japan

A palaeocorystid raninoid crab, a species of Joeranina.

Joeranina houssineaui[69]

Sp. nov

Valid

Van Bakel

Late Cretaceous (Cenomanian)

 France

A palaeocorystid decapod, a species of Joeranina.

Kellnerius[70]

Gen. et sp. nov

Valid

Santana et al.

Early Cretaceous (Albian)

Romualdo Formation

 Brazil

A palaemonid. The type species is Kellnerius jamacaruensis.

Keslingiella? teresae[71]

Sp. nov

Valid

Salas, Rustán & Sterren

Devonian

Punta Negra Formation

 Argentina
 South Africa?

An ostracod belonging to the group Podocopida and possibly to the family Quasillitidae.

Knebelia totoroi[72]

Sp. nov

Valid[73]

Audo et al.

Late Jurassic (late Kimmeridgian to early Tithonian)

 Germany

An eryonid, a species of Knebelia.

Koskobilius[15]

Gen. et sp. nov

Valid

Klompmaker

Early Cretaceous (late Albian)

Eguino Formation

 Spain

A crab belonging to the group Majoidea and the family Priscinachidae. The type species is Koskobilius postangustus.

Laeviprosopon crassum[15]

Sp. nov

Valid

Klompmaker

Early Cretaceous (late Albian)

Eguino Formation

 Spain

A crab belonging to the family Prosopidae, a species of Laeviprosopon.

Laeviprosopon edoi[15]

Sp. nov

Valid

Klompmaker

Early Cretaceous (late Albian)

Eguino Formation

 Spain

A crab belonging to the family Prosopidae, a species of Laeviprosopon.

Laeviprosopon hispanicum[15]

Sp. nov

Valid

Klompmaker

Early Cretaceous (late Albian)

Eguino Formation

 Spain

A crab belonging to the family Prosopidae, a species of Laeviprosopon.

Laeviprosopon planum[15]

Sp. nov

Valid

Klompmaker

Early Cretaceous (late Albian)

Eguino Formation

 Spain

A crab belonging to the family Prosopidae, a species of Laeviprosopon.

Langdaia bullabalvanyensis[17]

Sp. nov

Valid

Crasquin in Forel et al.

Early Triassic

 Hungary
 Italy

An ostracod, a species of Langdaia.

Lapazites trinodis[71]

Sp. nov

Valid

Salas, Rustán & Sterren

Devonian (Emsian)

Punta Negra Formation

 Argentina

An ostracod belonging to the group Palaeocopida and the superfamily Kloedenelloidea.

Latheticocarcinus italicus[74]

Sp. nov

Valid

De Angeli & Ceccon

Eocene (Ypresian)

 Italy

A crab belonging to the family Homolidae.

Lessinilambrus[36]

Gen. et sp. nov

Valid

Beschin, De Angeli & Zarantonello

Eocene

 Italy

A crab belonging to the family Parthenopidae. Genus includes new species L. paleogenicus.

Liuzhinia bankutensis[17]

Sp. nov

Valid

Forel in Forel et al.

Late Permian to Early Triassic

 Hungary

An ostracod, a species of Liuzhinia.

Liuzhinia venninae[17]

Sp. nov

Valid

Forel in Forel et al.

Early Triassic (Griesbachian)

Gerennavár Limestone Formation

 Hungary

An ostracod, a species of Liuzhinia.

Loxoconcha akrabouensis[34]

Sp. nov

Valid

Andreu et al.

Late Cretaceous

 Morocco

An ostracod.

Loxoconcha ? ngolothiensis[32]

Sp. nov

Valid

Sarr

Paleocene (Thanetian)

 Senegal

An ostracod belonging to the family Loxoconchidae.

Meridecaris[75]

Gen. et sp. nov

Valid

Stockar & Garassino

Middle Triassic (Ladinian)

Meride Limestone

  Switzerland

A clytiopsid decapod. The type species is Meridecaris ladinica.

Metahomarus[76]

Gen. et sp. nov

Valid

Franţescu

Early Cretaceous (Albian)

Pawpaw Shale

 United States

A lobster. The type species is Metahomarus reidi.

Mexicania[77]

Gen. et sp. nov

Valid

Garassino et al.

Late Cretaceous (Cenomanian)

Sierra Madre Formation

 Mexico

A member of Sicyoniidae. The type species is M. grijalvaensis.

Microcheilinella egerensis[17]

Sp. nov

Valid

Forel in Forel et al.

Early Triassic (Griesbachian)

Gerennavár Limestone Formation

 Hungary

A pachydomellid ostracod, a species of Microcheilinella.

Moianella[78]

Gen. et sp. nov

Valid

Ossó & Domínguez

Late Eocene (Priabonian)

 Spain

A crab, possibly a member of Cancroidea. The type species is Moianella cervantesi.

Mokaya[77]

Gen. et sp. nov

Valid

Garassino et al.

Late Cretaceous (Cenomanian)

Sierra Madre Formation

 Mexico

A member of Sergestidae. The type species is M. changoensis.

Mongolitubulus aspermachaera[79]

Sp. nov

Valid

Topper et al.

Cambrian (Furongian)

 Sweden

A mongolitubulid, a member of Bradoriida; a species of Mongolitubulus.

Nantocyclois[37]

Gen. et sp. nov

Valid

Beschin, Busulini & Tessier

Eocene

 Italy

A calappid crab. The type species is Nantocyclois eocenicus.

Navarrara[15]

Gen. et sp. nov

Valid

Klompmaker

Early Cretaceous (late Albian)

Eguino Formation

 Spain

A crab belonging to the group Glaessneropsoidea and the family Longodromitidae. The type species is Navarrara betsieae.

Nestoria donggouensis[60]

Sp. nov

Valid

Wang

Early Cretaceous

Qingshuihe Formation

 China

A clam shrimp, a species of Nestoria.

Nestoria jungarensis[60]

Sp. nov

Valid

Wang

Early Cretaceous

Qingshuihe Formation

 China

A clam shrimp, a species of Nestoria.

Nestoria shawanensis[60]

Sp. nov

Valid

Wang

Early Cretaceous

Qingshuihe Formation

 China

A clam shrimp, a species of Nestoria.

Newnhamia mckenziana[44]

Sp. nov

Valid[45]

Matzke-Karasz et al.

Early Miocene (23–16 Ma)

Riversleigh

 Australia

An ostracod, a species of Newnhamia.

Nigeroloxoconcha itanhaensis[25]

Sp. nov

Valid

Nicolaidis & Piovesan in Piovesan et al.

Late Cretaceous (Cenomanian)

Itanhaém Formation

 Brazil

An ostracod belonging to the family Loxoconchidae.

Notopoides nantoensis[37]

Sp. nov

Valid

Beschin, Busulini & Tessier

Eocene

 Italy

A raninid crab, a species of Notopoides.

Octoeurax[22]

Gen. et sp. nov

Valid

Robins, Feldmann & Schweitzer

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A munidopsid galatheoid. The type species is Octoeurax acaresprora.

Ordonyia dabadibensis[19]

Sp. nov

Valid

Boukhary & Sharabi in Boukhary et al.

Paleocene

 Egypt

An ostracod.

Orhomalus arpi[52]

Sp. nov

Valid

Schweigert et al.

Early Jurassic (late Toarcian)

 Germany

A hermit crab, a species of Orhomalus.

Orhomalus dubrullei[52]

Sp. nov

Valid

Schweigert et al.

Early Jurassic (late Pliensbachian)

 France

A hermit crab, a species of Orhomalus.

Orthonotacythere spinata[32]

Sp. nov

Valid

Sarr

Paleocene (Thanetian)

 Senegal

An ostracod belonging to the family Cytheruridae.

Oryctocaris[80]

Gen. et comb. nov

Valid[81]

Bergmann & Rust

Early Devonian

 Germany

A phyllocarid; a new genus for "Nahecaris" balssi Broili (1930).

Paguritergites[82]

Gen. et sp. nov

Valid

Fraaije et al.

Early Cretaceous (late Albian)

Eguino Formation

 Spain

A hermit crab, possibly a member of Paguridae. The type species is Paguritergites yvonnecooleae.

Palaeobenthesicymus[83]

gen. et comb nov

Valid

Audo & Charbonnier

Late Santonian

Sahel Alma Lagerstätte

 Lebanon

A benthesicymid penaeoid, a new genus for "Penaeus" libanensis (Brocchi, 1875).

Palaeocypridina[57]

Gen. et 2 sp. nov

Valid

Crasquin in Sebe, Crasquin & Grădinaru

Middle Triassic (Anisian)

 Romania

An ostracod belonging to the family Cypridinellidae. The type species is P. tulceaensis Sebe in Sebe, Crasquin & Grădinaru; genus also includes P. agighiolensis Sebe in Sebe, Crasquin & Grădinaru.

Palinurus veronicae[76]

Sp. nov

Valid

Franţescu

Early Cretaceous (Albian)

Pawpaw Shale

 United States

A lobster, a species of Palinurus.

Panopeus bericus[37]

Sp. nov

Valid

Beschin, Busulini & Tessier

Eocene

 Italy

A panopeid crab, a species of Panopeus.

Paracypria? elongata[39]

Sp. nov

Valid

Do Carmo et al.

Early Cretaceous (Aptian)

Alagamar Formation

 Brazil

An ostracod, possibly a species of Paracypria.

Paracypris eniotmetos[25]

Sp. nov

Valid

Nicolaidis & Piovesan in Piovesan et al.

Cretaceous (Albian-Santonian)

Florianópolis Formation
Itanhaém Formation
Juréia Formation
São Mateus Formation

 Brazil

An ostracod belonging to the family Candonidae.

Paraglyphea mistelgauensis[27]

Sp. nov

Valid

Charbonnier et al.

Early Jurassic

 Germany

A member of the family Glypheidae.

Parallelogastrus[76]

Gen. et sp. nov

Valid

Franţescu

Early Cretaceous (Albian)

Pawpaw Shale

 United States

A lobster. The type species is Parallelogastrus gabii.

Parapalaemonetes[23]

Gen. st sp. nov

Valid

Brandt & Schulz

Middle Triassic (Ladinian)

Meißner Formation

 Germany

A member of the family Palaemonidae. The type species is P. thuringiacus.

Parapylochelitergites[82]

Gen. et sp. nov

Valid

Fraaije et al.

Early Cretaceous (late Albian)

Eguino Formation

 Spain

A hermit crab, possibly a member of Parapylochelidae. The type species is Parapylochelitergites pustulosus.

Paratetralia sulcata[29]

Sp. nov

Valid

De Angeli & Ceccon

Eocene (Ypresian)

 Italy

A trapeziid trapezioid, a species of Paratetralia.

Parthenope chiampensis[36]

Sp. nov

Valid

Beschin, De Angeli & Zarantonello

Eocene

 Italy

A crab belonging to the family Parthenopidae.

Pauline[84]

Gen. et sp. nov

Valid

Siveter et al.

Silurian

Wenlock Group

 United Kingdom

A cylindroleberidid myodocope ostracod. The type species is Pauline avibella.

Pegomyrmekella[22]

Gen. et sp. nov

Valid

Robins, Feldmann & Schweitzer

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A munidopsid galatheoid. The type species is Pegomyrmekella chaulia.

Philyra karkata[85]

Sp. nov

Valid

Rai et al.

Early Miocene

Khari Nadi Formation

 India

A leucosiid crab, a species of Philyra.

Pircawayra[71]

Gen. et comb. et sp. nov

Valid

Salas, Rustán & Sterren

Devonian

Belén Formation
Punta Negra Formation
Talacasto Formation

 Argentina
 Bolivia
 South Africa?

An ostracod belonging to the group Palaeocopida and the superfamily Kloedenelloidea. The type species is "Pinnatulites"? peregrina Přibyl (1984); genus also includes new species P. gigantea.

Planocarcinus johnjagti[67]

Sp. nov

Valid

Bermúdez et al.

Early Cretaceous (late Aptian)

 Colombia

A necrocarcinid raninoid crab, a species of Planocarcinus.

Pokornyella lagandongi[32]

Sp. nov

Valid

Sarr

Paleocene (Thanetian)

 Senegal

An ostracod belonging to the family Hemicytheridae.

Polzicaris[86]

Gen. et comb. nov

Valid

Haug et al.

Late Cretaceous (Santonian)

 Lebanon

A member of Achelata; a new genus for "Eryoneicus" sahelalmae Roger (1944).

Pontocypris tadighoustensis[34]

Sp. nov

Valid

Andreu et al.

Late Cretaceous

 Morocco

An ostracod.

Poupelicaris[87]

Gen. et sp. nov

Valid

Racheboeuf & Gourvennec

Devonian (Lochkovian)

 Bolivia

A member of Phyllocarida. The type species is Poupelicaris elegans.

Praemonoleaia[40]

Gen. et 6 sp. nov

Valid

Niu in Niu et al.

Late Permian

Taohaiyingzi Formation

 China

A clam shrimp. Genus contains species Praemonoleaia ampulliformis, P. elliptica, P. oblonga, P. ovalis, P. rotunda and P. subtriangularis.

Procytherura? elongatissima[34]

Sp. nov

Valid

Andreu et al.

Late Cretaceous

 Morocco

An ostracod.

Protobuntonia baolensis[32]

Sp. nov

Valid

Sarr

Paleocene (Selandian-Thanetian)

 Senegal

An ostracod belonging to the family Trachyleberididae.

Puriana sokonensis[32]

Sp. nov

Valid

Sarr

Paleocene (Thanetian)

 Senegal

An ostracod belonging to the family Trachyleberididae.

Pylochelitergites alatus[82]

Sp. nov

Valid

Fraaije et al.

Early Cretaceous (late Albian)

Eguino Formation

 Spain

A hermit crab, possibly a member of Pylochelidae; a species of Pylochelitergites.

Pylochelitergites rugosus[82]

Sp. nov

Valid

Fraaije et al.

Early Cretaceous (late Albian)

Eguino Formation

 Spain

A hermit crab, possibly a member of Pylochelidae; a species of Pylochelitergites.

Pylochelitergites stramberkensis[33]

Sp. nov

Valid

Fraaije et al.

Late Jurassic to Early Cretaceous (Tithonian to early Berriasian)

Štramberk Limestone

 Czech Republic

A hermit crab, possibly a member of Pylochelidae; a species of Pylochelitergites.

Rathbunassa[67]

Gen. et comb. nov

Valid

Hyžný in Bermúdez et al.

Early Cretaceous

 Colombia
 United States

A callianassid decapod, a new genus for "Callianassa" aquilae Rathbun (1935).

Rectaglyphea[27]

Gen. et sp. nov

Valid

Charbonnier et al.

Cretaceous

 United Kingdom

A member of the family Glypheidae. Genus includes new species R. howardae.

Rehacythereis errachidiaensis[34]

Sp. nov

Valid

Andreu et al.

Late Cretaceous

 Morocco

An ostracod.

Rehacythereis zizensis[34]

Sp. nov

Valid

Andreu et al.

Late Cretaceous

 Morocco

An ostracod.

Reviya praecurukensis[17]

Sp. nov

Valid

Forel in Forel et al.

Late Permian (Changhsingian)

Nagyvisnyó Limestone Formation

 Hungary
 Serbia?

A kirkbyid ostracod, a species of Reviya.

Rolerithosia[88]

Gen. et sp. nov

Valid

Collins, Villier & Breton

Late Cretaceous (Cenomanian)

 France

An etyiid crab. The type species is Rolerithosia lobulata.

Rosacythere denticulata[89]

Sp. nov

Valid

Tibert et al.

Early Cretaceous

Escucha Formation

 Spain

An ostracod.

Rosagammarus[90]

Gen. et sp. nov

Valid

McMenamin, Zapata & Hussey

Late Triassic (Norian)

Luning Formation

 United States

Originally interpreted as an amphipod;[90] the holotype specimen was subsequently reinterpreted as the right half of a decapod tail of uncertain taxonomic affinity.[91][92] The type species is Rosagammarus minichiellus.

Scalpellum carentanensis[30]

Sp. nov

Valid

Carriol

Pliocene

 France

A barnacle, a species of Scalpellum.

Schernfeldia[93]

Gen. et sp. nov

Valid

Winkler

Late Jurassic (early Tithonian)

Eichstätt Formation

 Germany

An acratiid caridean shrimp of uncertain phylogenetic placement. The type species is Schernfeldia schweigerti.

Schizocythere wiedmanni[32]

Sp. nov

Valid

Sarr

Paleocene (Thanetian)

 Senegal

An ostracod belonging to the family Schizocytheridae.

Schobertella[52]

Gen. et 2 sp. nov

Valid

Schweigert et al.

Early Jurassic (late Hettangian to late Pliensbachian)

 Germany

A hermit crab. Genus contains species Schobertella simonsenetlangi and Schobertella hoelderi.

Scyllarella adinae[76]

Sp. nov

Valid

Franţescu

Early Cretaceous (Albian)

Pawpaw Shale

 United States

A lobster, a species of Scyllarella.

Seorsus kauffmani[47]

Sp. nov

Valid

Feldmann et al.

Late Cretaceous (Turonian)

Mancos Shale

 United States

A dakoticancroid crab, a species of Seorsus.

Serrablopluma[62]

Gen. et sp. nov

Valid

Artal et al.

Eocene

 Spain

A retroplumid crab. The type species is Serrablopluma diminuta.

Shemonaella? olempskaella[17]

Sp. nov

Valid

Forel in Forel et al.

Late Permian (Changhsingian)

Nagyvisnyó Limestone Formation

 Hungary

A paraparchitid ostracod, a possible species of Shemonaella.

Shiheziestheria[60]

Gen. et sp. nov

Valid

Wang

Early Cretaceous

Qingshuihe Formation

 China

A clam shrimp. The type species is Shiheziestheria pangi.

Shipingia weemsi[94]

Sp. nov

Valid

Kozur, Franz & Bachmann

Late Triassic (late Carnian to early Norian)

Arnstadt Formation
Weser Formation

 Germany

A clam shrimp, a species of Shipingia.

Soleryon[53]

Gen. et 2 sp. et comb. nov

Valid[54]

Audo et al.

Middle[95] and Late Jurassic

 France
 Germany

An eryonid. Genus contains two new species: Soleryon amicalis and S. schorri, as well as "Eryon" perroni Étallon (1859) and "Eryon" giganteus Van Straelen (1923).[95]

Spinogriphus[96]

Gen. et sp. nov

Valid

Jaume, Pinardo-Moya & Boxshall

Early Cretaceous (late Barremian)

 Spain

An acadiocaridid, a member of Spelaeogriphacea. The type species is Spinogriphus ibericus.

Squilla taulinanus[97]

Sp. nov

Valid

Ahyong, Charbonnier & Garassino

Miocene (Burdigalian)

 France

A squillid mantis shrimp, a species of Squilla.

Stenodactylina rogerfurzei[98]

Sp. nov

Valid

Schweigert

Middle Jurassic (late Aalenian)

 Germany

An erymid decapod, a species of Stenodactylina.

Systenobythere[99]

Gen. et sp. nov

Valid

Warne & Whatley

Late Miocene

 Australia

An ostracod. The type species is Systenobythere archboldi.

Tanidromites maerteni[100]

Sp. nov

Valid

Fraaije et al.

Middle Jurassic (Bajocian)

 France

A tanidromitid crab, a relative of homolodromiids; a species of Tanidromites. Announced in 2013; validated in 2023.[101]

Tetralia vicetina[29]

Sp. nov

Valid

De Angeli & Ceccon

Eocene (Ypresian)

 Italy

A tetraliid trapezioid, a species of Tetralia.

Theriosynoecum arinnoensis[89]

Sp. nov

Valid

Tibert et al.

Early Cretaceous

Escucha Formation

 Spain

An ostracod.

Theriosynoecum colini[39]

Sp. nov

Valid

Do Carmo et al.

Early Cretaceous (Aptian)

Alagamar Formation

 Brazil

An ostracod, a species of Theriosynoecum.

Theriosynoecum escuchaensis[89]

Sp. nov

Valid

Tibert et al.

Early Cretaceous

Escucha Formation

 Spain

An ostracod.

Theriosynoecum guzzoi[39]

Sp. nov

Valid

Do Carmo et al.

Early Cretaceous (Aptian)

Alagamar Formation

 Brazil

An ostracod, a species of Theriosynoecum.

Tzeltalpenaeus[77]

Gen. et sp. nov

Valid

Garassino et al.

Late Cretaceous (Cenomanian)

Sierra Madre Formation

 Mexico

A member of Penaeidae. The type species is T. exilichelatus.

Upogebia aronae[102]

Sp. nov

Valid

Haug, Nyborg & Vega

Middle Eocene

Santiago Formation

 United States

A member of Upogebiidae, a species of Upogebia.

Urocythereis castelaini[32]

Sp. nov

Valid

Sarr

Paleocene (Thanetian)

 Senegal

An ostracod belonging to the family Hemicytheridae.

Urocythereis sinesaloumensis[32]

Sp. nov

Valid

Sarr

Paleocene-Eocene (Thanetian-Ypresian)

 Senegal

An ostracod belonging to the family Hemicytheridae.

Uromuellerina guiraudi[32]

Sp. nov

Valid

Sarr

Eocene (Ypresian-Lutetian)

 Senegal

An ostracod belonging to the family Hemicytheridae.

Veenia (Nigeria) mediacostarobusta[34]

Sp. nov

Valid

Andreu et al.

Late Cretaceous

 Morocco

An ostracod.

Veenia (Nigeria) tardaensis[34]

Sp. nov

Valid

Andreu et al.

Late Cretaceous

 Morocco

An ostracod.

Vetoplautus[22]

Gen. et sp. nov

Valid

Robins, Feldmann & Schweitzer

Late Jurassic (Tithonian)

Ernstbrunn Limestone

 Austria

A munidopsid galatheoid. The type species is Vetoplautus latimarginus.

Wrangelleryon[103]

Gen. et sp. nov

Valid

Feldmann, Schweitzer & Haggart

Early Jurassic (Hettangian)

Sandilands Formation

 Canada

An eryonid. The type species is Wrangelleryon perates.

Xestoleberis circinatus[34]

Sp. nov

Valid

Andreu et al.

Late Cretaceous

 Morocco

An ostracod.

Xestoleberis? preafricanensis[34]

Sp. nov

Valid

Andreu et al.

Late Cretaceous

 Morocco

An ostracod.

Zoquepenaeus[77]

Gen. et sp. nov

Valid

Garassino et al.

Late Cretaceous (Cenomanian)

Sierra Madre Formation

 Mexico

A member of Penaeidae. The type species is Z. spinirostratus.

Zygastrocarcinus carolinasensis[104]

Sp. nov

Valid

Klompmaker, Flores-Ventura & Vega

Late Cretaceous (late Campanian)

Cerro del Pueblo Formation

 Mexico

A homolid crab, a species of Zygastrocarcinus.

Insects

[edit]

Trilobites

[edit]
Name Novelty Status Authors Age Unit Location Notes Images

Acanthopyge (Belenopyge) estevei[105]

Sp. nov

Valid

Corbacho & Kier

Devonian (Pragian)

Ihandar Formation

 Morocco

A member of Lichida belonging to the family Lichidae, a species of Acanthopyge.

Acastava? lerougei[106]

Sp. nov

Valid

Van Viersen

Devonian

Riviere Formation

 Belgium

Adelogonus oblongus[107]

Sp. nov

Valid

Lin et al.

Cambrian

Liudaogou Group

 China

A polymerid trilobite belonging to the family Proasaphiscidae, a species of Adelogonus.

Akantharges mbareki[108]

Sp. nov

Valid

Corbacho & López-Soriano

Devonian (Eifelian)

 Morocco

A member of Lichida belonging to the family Lichidae, a species of Akantharges.

Basseiarges[108]

Gen. et sp. nov

Disputed

Corbacho & López-Soriano

Devonian (Eifelian)

 Morocco

A member of Lichida belonging to the family Lichidae. The type species is Basseiarges mellishae. Van Viersen (2021) considered Basseiarges to be a junior synonym of the genus Akantharges.[109]

Bathyurellus diclementsae[110]

Sp. nov

Valid

Fortey & Bruton

Early Ordovician

Kirtonryggen Formation

 Norway

A bathyurid proetid, a species of Bathyurellus.

Bolbocephalus gunnari[110]

Sp. nov

Valid

Fortey & Bruton

Early Ordovician

Kirtonryggen Formation

 Norway

A bathyurid proetid, a species of Bolbocephalus.

Bolbochasmops kaskovoensis[111]

Sp. nov

Valid

Krueger

Ordovician

 Russia

A member of the family Pterygometopidae.

Cambrunicornia? jafnaensis[112]

Sp. nov

Valid

Elicki & Geyer

Cambrian

 Jordan

A protolenine ellipsocephalid, possibly a species of Cambrunicornia.

Canningbole[113]

Gen. et 3 sp. nov

Valid

Feist & McNamara

Late Devonian

Virgin Hills Formation

 Australia

A member of Proetida. The type species is Canningbole latimargo; genus also contains C. henwoodorum and C. macromma.

Catochia hinlopensis[110]

Sp. nov

Valid

Fortey & Bruton

Early Ordovician

Kirtonryggen Formation

 Norway

A bathyurid proetid, a species of Catochia.

Ceraurinus zhuzhaiensis[114]

sp nov

Valid

Lee

Late Ordovician

Xiazhen Formation

 China

A cheirurid, a species of Ceraurinus.

Chapmanopyge[110]

Nom. nov

Valid

Fortey & Bruton

Ordovician

 Greenland
 Norway
 United States

A bathyurid proetid; a replacement name for Chapmania Loch (2007).

Chattiaspis? budili[115]

Sp. nov

Valid

Corbacho & López-Soriano

Ordovician (early Katian)

Ktaoua Formation

 Morocco

A member of Dalmanitidae, possibly a species of Chattiaspis.

Chlupaciparia (Australoparia)[113]

Subgen. et 2 sp. nov

Valid

Feist & McNamara

Late Devonian

Virgin Hills Formation

 Australia

A member of Proetida, a subgenus of Chlupaciparia. The type species is Chlupaciparia (Australoparia) australis; genus also contains C. (Australoparia) lata.

Chlupaciparia (Chlupaciparia) planiops[113]

Sp. nov

Valid

Feist & McNamara

Late Devonian

Virgin Hills Formation

 Australia

A member of Proetida, a species of Chlupaciparia.

Cirriticeps[116]

Gen. et sp. et comb. nov

Valid

Holloway

Silurian

 Australia

A member of Proetida belonging to the family Tropidocoryphidae. The type species is C. auranticum; genus also includes "Proetus" rattei Etheridge & Mitchell (1891).

Dianops algeriensis[117]

Sp. nov

Valid

Crônier et al.

Devonian (Famennian)

‘Argiles de Marhouma’ Formation

 Algeria

A member of Phacopidae.

Eurysymphysurina[110]

Gen. et sp. nov

Valid

Fortey & Bruton

Early Ordovician

Kirtonryggen Formation

 Norway

A symphysurinid cyclopygoid member of Asaphida. The type species is Eurysymphysurina spora.

Gabriellus kierorum[118]

Sp. nov

Valid

Corbacho & López-Soriano

Early Cambrian

Rosella Formation

 Canada

A member of Redlichiida belonging to the family Judomiidae, a species of Gabriellus.

Gandlops[119]

Gen. et comb. nov

Valid[120]

Bignon & Crônier

Devonian (Emsian)

 France
 Morocco
 Spain

An asteropygine phacopidan, a new genus for "Kayserops" brevispinosus Gandl (1972). Genus also contains Gandlops ogivalis (Morzadec, 1976), G. palenciae (Smeenk, 1983) and G. tamnrherta (Chatterton et al., 2006).

Harlandaspis[110]

Gen. et sp. nov

Valid

Fortey & Bruton

Early Ordovician

Kirtonryggen Formation

 Norway

A bathyurid proetid. The type species is Harlandaspis elongata.

Heliopyge sohensis[121]

Sp. nov

Valid

Ghobadi Pour in Ghobadi Pour et al.

Devonian (Frasnian)

 Iran

A member of Phacopida belonging to the family Acastidae.

Hibbertia aodiensis[114]

sp nov

Valid

Lee

Late Ordovician

Xiazhen Formation

 China

A harpetid, a species of Hibbertia.

Illaenus (Parillaenus) primoticus[110]

Sp. nov

Valid

Fortey & Bruton

Early Ordovician

Kirtonryggen Formation

 Norway

An illaenid corynexochid, a species of Illaenus.

Issalia[112]

Gen. et sp. nov

Valid

Elicki & Geyer

Cambrian

 Jordan

An ellipsocephaline ellipsocephalid. The type species is Issalia scutalis.

Jaylopygiella debruni[106]

Sp. nov

Valid

Van Viersen

Devonian

Riviere Formation

 Belgium

A member of Acastidae.

Jeffersonia striagena[110]

Sp. nov

Valid

Fortey & Bruton

Early Ordovician

Kirtonryggen Formation

 Norway

A bathyurid proetid, a species of Jeffersonia.

Jeffersonia viator[110]

Sp. nov

Valid

Fortey & Bruton

Early Ordovician

Kirtonryggen Formation

 Norway

A bathyurid proetid, a species of Jeffersonia.

Lachnostoma platypyga[110]

Sp. nov

Valid

Fortey & Bruton

Early Ordovician

Kirtonryggen Formation

 Norway

An asaphid, a species of Lachnostoma.

Leiostegium spongiosum[110]

Sp. nov

Valid

Fortey & Bruton

Early Ordovician

Kirtonryggen Formation

 Norway

A leiostegiid corynexochid, a species of Leiostegium.

Linguaphillipsia simekuhensis[122]

Sp. nov

Valid

Hahn, Müller & Aghababalou

Carboniferous (Tournaisian)

Mobarak Formation

 Iran

A member of Phillipsiidae, a species of Linguaphillipsia.

Meteoraspis qinghaiensis[107]

Sp. nov

Valid

Lin et al.

Cambrian

Liudaogou Group

 China

A polymerid trilobite belonging to the family Crepicephalidae, a species of Meteoraspis.

Minicryphaeus[119]

Gen. et comb. nov

Valid[120]

Bignon & Crônier

Devonian (Pragian)

 Morocco

An asteropygine phacopidan, a new genus for "Pseudocryphaeus" minimus Morzadec (2001). Genus also contains Minicryphaeus quarterspinosus (Morzadec, 2001) and M. sarirus (Morzadec, 2001)

Morzadecops[119]

Gen. et comb. nov

Disputed

Bignon & Crônier

Devonian (middle Pragian to early Emsian)

 Morocco
 Spain

An asteropygine phacopidan, a new genus for "Pilletina" zguidensis Morzadec (2001). Genus also contains Morzadecops matutina (Gandl, 1972). The genus Morzadecops was considered to be a junior synonym of the genus Feruminops by Van Viersen & Kloc (2022).[123]

Myopsolenites hyperion[112]

Sp. nov

Valid

Elicki & Geyer

Cambrian

 Jordan

A bathynotid, a species of Myopsolenites.

Pagetides johannis[124]

Sp. nov

Valid

Fletcher & Greene

Cambrian

Young's Cove Formation

 Canada

A species of Pagetides.

Palpebralia initialis[113]

Sp. nov

Valid

Feist & McNamara

Late Devonian

Virgin Hills Formation

 Australia

A member of Proetida, a species of Palpebralia.

Palpebralia pustulata[113]

Sp. nov

Valid

Feist & McNamara

Late Devonian

Virgin Hills Formation

 Australia

A member of Proetida, a species of Palpebralia.

Palpebralina[113]

Gen. et 3 sp. et comb. nov

Valid

Feist & McNamara

Late Devonian

 Australia
 France

A member of Proetida. The type species is Palpebralina pseudopalpebralis; genus also contains new species P. minor and P. ocellifer, as well as "Palpebralia" palpebralis globoculata Feist in Feist and Schindler, 1994 (raised by Feist and McNamara to the rank of a separate species Palpebralina globoculata) and "Palpebralia" latepalpebralis Feist in Feist and Schindler (1994).

Paranomocarella dingmaoshanensis[107]

Sp. nov

Valid

Lin et al.

Cambrian

Liudaogou Group

 China

A polymerid trilobite belonging to the family Anomocarellidae, a species of Paranomocarella.

Parvilichas[125]

Gen. et sp. nov

Valid

Corbacho & Vela

Ordovician (Floian)

Fezouata Formation

 Morocco

A member of Lichida belonging to the family Lichidae. The type species is Parvilichas marochii.

Peltabellia glabra[110]

Sp. nov

Valid

Fortey & Bruton

Early Ordovician

Kirtonryggen Formation

 Norway

A bathyurid proetid, a species of Peltabellia.

Pennarbedops[119]

Gen. et comb. nov

Valid[120]

Bignon & Crônier

Devonian (late Emsian)

 France
 Spain

An asteropygine phacopidan, a new genus for "Rhenops" babini Morzadec (1983). Genus also contains Pennarbedops circumapodemus (Smeenk, 1983). The genus Pennardbedops was considered to be a junior synonym of the genus Hollardops by Van Viersen & Kloc (2022).[123]

Phacops tamtertensis[117]

Sp. nov

Valid

Crônier et al.

Devonian (Famennian)

‘Argiles de Marhouma’ Formation

 Algeria

A member of Phacopidae.

Phacops zeregensis[117]

Sp. nov

Valid

Crônier et al.

Devonian (Famennian)

‘Argiles de Marhouma’ Formation

 Algeria

A member of Phacopidae.

Phaseolops? bobowensi[110]

Sp. nov

Valid

Fortey & Bruton

Early Ordovician

Kirtonryggen Formation

 Norway

A member of Proetidae, possibly a species of Phaseolops.

Placosema qinghaiense[107]

Sp. nov

Valid

Lin et al.

Cambrian

Liudaogou Group

 China

A polymerid trilobite belonging to the family Catillicephalidae, a species of Placosema.

Pliomerina tashanensis[114]

sp nov

Valid

Lee

Late Ordovician

Xiazhen Formation

 China

A pliomerid, a species of Pliomerina.

Pteroparia extrema[113]

Sp. nov

Valid

Feist & McNamara

Late Devonian

Virgin Hills Formation

 Australia

A member of Proetida, a species of Pteroparia.

Punka latissima[110]

Sp. nov

Valid

Fortey & Bruton

Early Ordovician

Kirtonryggen Formation

 Norway

A bathyurid proetid, a species of Punka.

Raymondaspis? pingpong[110]

Sp. nov

Valid

Fortey & Bruton

Early Ordovician

Kirtonryggen Formation

 Norway

A styginid corynexochid, possibly a species of Raymondaspis.

Remopleurides xiazhenensis[114]

sp nov

Valid

Lee

Late Ordovician

Xiazhen Formation

 China

A remopleurid, a species of Remopleurides.

Rhenogriffides alborzensis[122]

Sp. nov

Valid

Hahn, Müller & Aghababalou

Carboniferous (Tournaisian)

Mobarak Formation

 Iran

A member of Phillipsiidae, a species of Rhenogriffides.

Rudybole[113]

Gen. et comb. et sp. nov

Valid

Feist & McNamara

Late Devonian

 Australia
 France

A member of Proetida. A new genus for "Drevermannia" brecciae Richter (1913); genus also contains "Drevermannia" adorfensis Richter (1913), "Drevermannia" nodannulata Richter (1913), "Palpebralia" nodannulata concava Feist in Feist and Schindler, 1994 (raised by Feist and McNamara to the rank of a separate species Rudybole concava) and a new species Rudybole depressa.

Scopelochasmops mukranensis[111]

Sp. nov

Valid

Krueger

Ordovician

 Germany

A member of the family Pterygometopidae.

Scopelochasmops praewrangeli[111]

Sp. nov

Valid

Krueger

Ordovician

Europe

A member of the family Pterygometopidae.

Shabanovia[126]

Nom. nov

Valid

Doweld

Middle Cambrian

 Russia

An alokistocarid; a replacement name for Furia Egorova and Shabanov (1987).

Svalbardicurus[110]

Gen. et sp. nov

Valid

Fortey & Bruton

Early Ordovician

Kirtonryggen Formation

 Norway

A hystricurid aulacopleurid. The type species is Svalbardicurus delicatus.

Tangshanaspis silveri[127]

Sp. nov

Valid

Westrop

Late Cambrian (Furongian)

Signal Mountain Formation

 United States

A member of Missisquoiidae, a species of Tangshanaspis.

Tayanaspis[112]

Gen. et sp. nov

Valid

Elicki & Geyer

Cambrian

 Jordan

An ellipsocephaline ellipsocephalid. The type species is Tayanaspis bulbosus.

Toxochasmops (Toxochasmops) luebzensis[111]

Sp. nov

Valid

Krueger

Ordovician

Europe

A member of the family Pterygometopidae.

Toxochasmops (Toxochasmops) praewesenbergensis[111]

Sp. nov

Valid

Krueger

Ordovician

Europe

A member of the family Pterygometopidae.

Trimerocephalus chopini[128]

Sp nov

Valid

Kin & Błażejowski

Late Devonian (Early Famennian)

 Poland

A phacopid, a species of Trimerocephalus.

Uhaymiria[112]

Gen. et sp. nov

Valid

Elicki & Geyer

Cambrian

 Jordan

An ellipsocephaline ellipsocephalid. The type species is Uhaymiria glabra.

Uromystrum drepanon[110]

Sp. nov

Valid

Fortey & Bruton

Early Ordovician

Kirtonryggen Formation

 Norway

A bathyurid proetid, a species of Uromystrum.

Valdariops porkuniensis[111]

Sp. nov

Valid

Krueger

Ordovician

Europe

A member of the family Pterygometopidae.

Valdariops postnybiensis[111]

Sp. nov

Valid

Krueger

Ordovician

Europe

A member of the family Pterygometopidae.

Valdariops ruegensis[111]

Sp. nov

Valid

Krueger

Ordovician

Europe

A member of the family Pterygometopidae.

Valdariops scheffelae[111]

Sp. nov

Valid

Krueger

Ordovician

 Germany

A member of the family Pterygometopidae.

Valdariops schneideri[111]

Sp. nov

Valid

Krueger

Ordovician

Europe

A member of the family Pterygometopidae.

Valdariops vierradensis[111]

Sp. nov

Valid

Krueger

Ordovician

Europe

A member of the family Pterygometopidae.

Vietnamia yushanensis[114]

sp nov

Valid

Lee

Late Ordovician

Xiazhen Formation

 China

A calymenid, a species of Vietnamia.

Zhifangia damghanensis[122]

Sp. nov

Valid

Hahn, Müller & Aghababalou

Carboniferous (Tournaisian)

Mobarak Formation

 Iran

A member of Proetidae. Originally described as a species of Zhifangia;[122] subsequently transferred to the genus Pudoproetus by Müller & Hahn (2015).[129]

Others

[edit]
Name Novelty Status Authors Age Unit Location Notes Images

Adelophthalmus piussii[130]

Sp. nov

Valid

Lamsdell, Simonetto & Selden

Late Carboniferous

 Italy

A eurypterid, a species of Adelophthalmus.

Aglaspella sanduensis[131]

Sp. nov

Valid

Lerosey-Aubril, Ortega-Hernández & Zhu

Cambrian (Furongian)

Sandu Formation

 China

A member of Aglaspidida, a species of Aglaspella.

Arthroaspis[132]

Gen. et sp. nov

Valid

Stein et al.

Cambrian (Montezuman)

Sirius Passet Lagerstätte

 Greenland

An arthropod of uncertain phylogenetic placement, probably a relative of trilobites. The type species is Arthroaspis bergstroemi.

Austromarrella[133]

Gen. et sp.

Valid

Haug et al.

Cambrian (Late Templetonian to Early Floran)

Monastery Creek Phosphorite Formation

 Australia

A marrellomorph. The type species is Austromarrella klausmuelleri.

Dvulikiaspis[134]

Gen. et comb. nov

Valid[135]

Marshall et al.

Early Devonian (Lochkovian)

 Russia

A member of Chasmataspidida, a new genus for "Tylopterella" menneri (initially thought to be a eurypterid).

Hesselbonia[131]

Gen. et comb. nov

Valid

Lerosey-Aubril, Ortega-Hernández & Zhu

Late Cambrian

St Lawrence Formation

 United States

A member of Aglaspidida, a new genus for "Aglaspella" eatoni.

Kachinophilus[136]

Gen. et sp. nov

Valid[137]

Bonato, Edgecombe & Minelli

Late Cretaceous (earliest Cenomanian)

 Myanmar

A geophilomorph centipede found in Burmese amber. The type species is Kachinophilus pereirai.

Kasibelinurus yueya[138]

Sp. nov

Valid

Lamsdell, Xue & Selden

Early Devonian (Lochkovian)

Xiaxishancun Formation

 China

Originally classified as a kasibelinurid xiphosuran and a species of Kasibelinurus;[138] Selden, Lamsdell & Qi (2015) reinterpreted it as a chelicerate belonging to the lineage that branched from the main euchelicerate lineage prior to the divergence of Eurypterida, Arachnida and Chasmataspidida and made it the type species of a separate genus Houia.[139]

Kootenichela[140]

Gen. et sp. nov

Valid

Legg

Middle Cambrian

Stephen Formation

 Canada

A member of paraphyletic "Megacheira" closely related to Worthenella. The type species is Kootenichela deppi. Later study considered this species to be chimera.[141]

Loricicaris[142]

Gen. et sp. nov

Valid[143]

Legg & Caron

Cambrian

Burgess Shale

 Canada

A basal, bivalved arthropod. The type species is Loricicaris spinocaudatus.

Nahlyostaspis[134]

Gen. et sp. nov

Valid[135]

Marshall et al.

Early Devonian (Lochkovian)

 Russia

A member of Chasmataspidida. The type species is Nahlyostaspis bergstroemi.

Nereocaris briggsi[142]

Sp. nov

Valid[143]

Legg & Caron

Cambrian

Burgess Shale

 Canada

A basal, bivalved arthropod; a species of Nereocaris.

Norikeya[144]

Gen. et sp. nov

Valid

Tanaka, Siveter & Williams

Early Devonian

 Japan

A bivalved arthropod of uncertain phylogenetic placement. The type species is Norikeya onoi.

Palaeomarachne[145]

Gen. et sp. nov

Valid

Rudkin et al.

Late Ordovician

William Lake Konservat-Lagerstätte

 Canada

A sea spider. The type species is Palaeomarachne granulata.

Paraeurypterus[146]

Gen. et sp. nov

Valid

Lamsdell, Hoşgör & Selden

Upper Ordovician

Şort Tepe Formation

 Turkey

A eurypterid. The type species is Paraeurypterus anatoliensis.

Parastemmiulus[147]

Gen. et sp. nov

Valid

Riquelme et al.

Miocene

 Mexico

A stemmiulid millipede. The type species is Parastemmiulus elektron.

Pentapantopus[148]

Gen. et sp. nov

Valid

Kühl, Poschmann & Rust

Early Devonian

 Germany

A sea spider. The type species is Pentapantopus vogteli.

Pentlandopterus[146]

Gen. et comb. nov

Valid

Lamsdell, Hoşgör & Selden

Silurian

 United Kingdom

A eurypterid; a new genus for "Eurypterus" minor Laurie (1899).

Sinoleperditia (Sinoleperditia) tongvaiensis[149]

Sp. nov

Valid

Wang

Early Devonian

Khao Loc Formation

 Vietnam

A member of Leperditicopida (a group of arthropods of uncertain affinities, classified as ostracods or excluded from this group by different authors), a species of Sinoleperditia.

Skrytyaspis[134]

Gen. et sp. nov

Valid[135]

Marshall et al.

Early Devonian (Lochkovian)

 Russia

A member of Chasmataspidida. The type species is Skrytyaspis andersoni.

Strobilopterus proteus[150]

Sp. nov

Valid

Lamsdell & Selden

Devonian (Pragian)

Beartooth Butte Formation

 United States

A eurypterid, a species of Strobilopterus.

Tremaglaspis vanroyi[151]

Sp. nov

Valid

Lerosey-Aubril et al.

Cambrian (Guzhangian)

Weeks Formation

 United States

A member of Aglaspidida, a species of Tremaglaspis.

References

[edit]
  1. ^ Ladislav Miko; Jan Mourek; Ioana N. Meleg; Oana T. Moldovan (2013). "Oribatid mite fossils from pre-Quaternary sediments in Slovenian caves II. Amiracarus pliocennatus n.gen., n.sp. (Microzetidae) from Pliocene, with comments on the other species of the genus". Zootaxa. 3670 (4): 557–578. doi:10.11646/zootaxa.3670.4.8. PMID 26438960. S2CID 38910961.
  2. ^ Wilson R. Lourenço (2013). "A new species of Chaerilobuthus Lourenço & Beigel, 2011 from Cretaceous Burmese amber (Scorpiones: Chaerilobuthidae)". Acta Biológica Paranaense. 42 (1–4): 1–5. doi:10.5380/abpr.v42i1-4.35095.
  3. ^ Fabio M. Dalla Vecchia; Paul A. Selden (2013). "A Triassic spider from Italy". Acta Palaeontologica Polonica. 58 (2): 325–330. doi:10.4202/app.2011.0132. hdl:1808/17084.
  4. ^ Robert W. Gess (2013). "The earliest record of terrestrial animals in Gondwana: A scorpion from the Famennian (Late Devonian) Witpoort Formation of South Africa". African Invertebrates. 54 (2): 373–379. Bibcode:2013AfrIn..54..373G. doi:10.5733/afin.054.0206.
  5. ^ a b Ekaterina Sidorchuk; Michel Bertrand (2013). "New fossil labidostomatids (Acari: Labidostomatidae) from Eocene amber and presence of an apustulate species in Europe". Acarologia. 53 (1): 25–39. doi:10.1051/acarologia/20132079.
  6. ^ a b c Ricardo Pérez-de la Fuente; Erin E. Saupe; Paul A. Selden (2013). "New lagonomegopid spiders (Araneae: †Lagonomegopidae) from Early Cretaceous Spanish amber". Journal of Systematic Palaeontology. 11 (5): 531–553. doi:10.1080/14772019.2012.725679. S2CID 55375767.
  7. ^ a b c "ZooBank.org".
  8. ^ Selden, P. A.; Shih, C.K.; Ren, D. (2013). "A giant spider from the Jurassic of China reveals greater diversity of the orbicularian stem group". Naturwissenschaften. 100 (12): 1171–1181. Bibcode:2013NW....100.1171S. doi:10.1007/s00114-013-1121-7. PMC 3889289. PMID 24317464.
  9. ^ Jason A. Dunlop; Ronny Rößler (2013). "The youngest trigonotarbid Permotarbus schuberti n. gen., n. sp. from the Permian Petrified Forest of Chemnitz in Germany". Fossil Record. 16 (2): 229–243. doi:10.1002/mmng.201300012.
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