Xenophora

Xenophora; Temporal range: Late Cretaceous–Recent PreꞒ Ꞓ O S D C P T J K Pg N
	A ventral view of a shell of Xenophora pallidula, with attached gastropod shells
	A lateral view of the same shell
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Caenogastropoda
Order:	Littorinimorpha
Family:	Xenophoridae
Genus:	Xenophora; Fischer von Waldheim, 1807
Species
	See text
Synonyms
	Phorus Montfort, 1810; Xenophora (Austrophora) Kreipl, Alf & Kronenberg, 1999· accepted, alternate representation; † Xenophora (Endoptygma) Gabb, 1877 · accepted, alternate representation; Xenophora (Xenophora) Fischer von Waldheim, 1807· accepted, alternate representation; Xenophorus (incorrect subsequent spelling of Xenophora Fischer von Waldheim, 1807);

Xenophora, commonly called carrier shells, is a genus of medium-sized to large sea snails, marine gastropod mollusks in the family Xenophoridae, the carrier snails or carrier shells.^[2] The genus Xenophora is the type genus of the family Xenophoridae.

Etymology

The name Xenophora comes from two ancient Greek words, and means "bearing foreigners", so-called because in most species the snail cements pieces of rock or shells to its own shell at regular intervals as the shell grows.^[3]

Description

The shells of species within this genus vary from small to large (diameter of base without attachments 19–90 mm; height of shell 21–60 mm), depressed-conical, with narrow to very narrow, simple peripheral edge, non-porcellanous ventrally. Foreign objects are attached to all whorls, with generally more than 30% of dorsal surface obscured by these objects. The foreign objects are usually medium-sized to large.^[4] Although the foreign objects are usually mollusk shells, pebbles, or small pieces of coral rock, in some instances a bottle cap has been attached by the snail to its shell.

Species

The genus Xenophora includes the following species and subspecies:^[4]^[5]^[6]

Xenophora minuta Qi & Ma, 1986
Xenophora robusta Verrill, 1870

Subgenus Xenophora (Austrophora) Kreipl, Alf & Kronenberg, 1999

Xenophora flindersi (Cotton & Godfrey, 1938)

Subgenus Xenophora (Xenophora) Fischer von Waldheim, 1807

Xenophora cerea (Reeve, 1845)
Xenophora chinensis Philippi, 1841) ^{[citation needed]}
Xenophora conchyliophora (Born, 1780) - American carriersnail (type species of Xenophora)
Xenophora corrugata (Reeve, 1842)
Xenophora crispa (König, 1825)
Xenophora granulosa Ponder, 1983
Xenophora japonica Kuroda & Habe in Kuroda et al., 1971
Xenophora konoi Habe, 1953
Xenophora mekranensis (Newton, 1905)
Xenophora neozelanica Suter, 1908 - New Zealand carriershell
- Xenophora neozelandica neozelandica Suter, 1908
- Xenophora neozelandica kermadecensis Ponder, 1983
Xenophora pallidula (Reeve, 1842) - pallid carrier shell
Xenophora peroniana (Iredale, 1929)
- Xenophora peroniana peroniana (Iredale, 1929)
- Xenophora peroniana kondoi Ponder 1983
Xenophora senegalensis Fischer, 1873
Xenophora sinensis Qi & Li, 1986
Xenophora solarioides (Reeve, 1845)
Xenophora tenuis Fulton, 1938

Species brought into synonymy

Subgenus Xenophora (Stellaria) Möller, 1832: synonym of Stellaria Möller, 1832
Xenophora (Stellaria) testigera (Bronn, 1831): synonym of Stellaria testigera (Bronn, 1831)
Xenophora australis Souverbie & Montrouzier, 1870: synonym of Xenophora (Xenophora) solarioides (Reeve, 1845)
Xenophora calculifera (Reeve, 1843): synonym of Stellaria chinensis (Philippi, 1841)
Xenophora caperata sensu Petit de la Saussaye, 1857: synonym of Xenophora (Xenophora) senegalensis Fischer, 1873
Xenophora caribaea Petit de la Saussaye, 1857 - Caribbean carriersnail: synonym of Onustus caribaeus (Petit de la Saussaye, 1857)
Xenophora cavelieri Rochebrune, 1883: synonym of Xenophora (Xenophora) senegalensis Fischer, 1873
Xenophora digitata Martens, 1878: synonym of Stellaria testigera digitata Martens, 1878
Xenophora gigantea (Schepman, 1909): synonym of Stellaria gigantea (Schepman, 1909)
Xenophora helvacea Philippi, 1851: synonym of Onustus indicus (Gmelin, 1791)
Xenophora indica (Gmelin, 1791): synonym of Onustus indicus (Gmelin, 1791)
Xenophora japonica Kuroda & Habe, 1971: synonym of Xenophora (Xenophora) japonica Kuroda & Habe, 1971
Xenophora konoi Habe, 1953: synonym of Xenophora (Xenophora) mekranensis konoi Habe, 1953
Xenophora laevigata Fischer von Waldheim, 1807: synonym of Xenophora (Xenophora) conchyliophora (Born, 1780)
Xenophora lamberti Souverbie, 1871: synonym of Stellaria lamberti (Souverbie, 1871)
Xenophora longleyi Bartsch, 1931 - shingled carriersnail: synonym of Onustus longleyi Bartsch, 1931
Xenophora meandrina Fischer von Waldheim, 1807: synonym of Xenophora (Xenophora) conchyliophora (Born, 1780)
Xenophora mediterranea Tiberi, 1863: synonym of Xenophora (Xenophora) crispa (König, 1825)
Xenophora neozelanica Suter, 1908: synonym of Xenophora (Xenophora) neozelanica neozelanica Suter, 1908
Xenophora regularis Habe & Okutani, 1983 : synonym of Xenophora (Xenophora) granulosa Ponder, 1983
Xenophora tenuis Hirase, 1934: synonym of Xenophora (Xenophora) tenuis Fulton, 1983
Xenophora torrida Kuroda & Ito, 1961 : synonym of Xenophora (Xenophora) cerea (Reeve, 1845)
Xenophora tricostata Fischer von Waldheim, 1807: synonym of Xenophora (Xenophora) conchyliophora (Born, 1780)
Xenophora tulearensis Stewart & Kosuge, 1993: synonym of Xenophora (Xenophora) corrugata (Reeve, 1842)
Xenophora vulcanica Fischer von Waldheim, 1807: synonym of Xenophora (Xenophora) conchyliophora (Born, 1780)
Xenophora wagneri Philippi, 1855: synonym of Onustus indicus (Gmelin, 1791)

Extinct species

Extinct species within this genus include:^[7]

†Xenophora carditigera Nielsen and DeVries 2002
†Xenophora cummulans Brongniart 1823
†Xenophora delecta Guppy 1876
†Xenophora deshayesi Michelotti 1847
†Xenophora eocenica Abbass 1967
†Xenophora floridana Mansfield 1930
†Xenophora infundibulum (Brocchi, 1814)^[8]
†Xenophora palaeoafra Gliozzi and Malatesta 1983
†Xenophora paulinae Nielsen and DeVries 2002
†Xenophora rugata Abbass 1967
†Xenophora tatei Harris 1897
†Xenophora terpstrai Dey 1961
†Xenophora textilina Dall 1892

Fossil record

Fossils of Xenophora are found in marine strata from the Cretaceous to Quaternary (age range: from 89.3 to 0.012 million years ago.). Fossils are known all over the world.^[7]^[9]

References

^ Fischer von Waldheim, Gotthelf (1807). Muséum Demidoff, ou catalogue systématique et raisonné des curiosités de la nature et de l'art. Vol. 3: Végétaux et animaux. p. 213.
^ WoRMS (2012). Xenophora. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=138664 on 2012-06-28
^ Xenophora.org website [1] accessed 24 April 2014
^ ^a ^b Kreipl, K. & Alf, A. (1999): Recent Xenophoridae. 148 pp. incl. 28 color plts. ConchBooks, Hackenheim, ISBN 3-925919-26-0.
^ Powell A. W. B. (1979). New Zealand Mollusca, William Collins Publishers Ltd, Auckland, New Zealand. ISBN 0-00-216906-1
^ OBIS Indo-Pacific Molluscan Database
^ ^a ^b Paleobiology Database
^ Stromboidea
^ G. Manganelli, V. Spadini, S. Cianfanelli The xenophorid gastropods of the Mediterranean Pliocene: the record of the Siena Basin^[usurped]

Kreipl K. & Alf A. (1999). Recent Xenophoridae. Conchbooks, Hackenheim > Germany. 148pp.

External links

Ladd, H.S. (1977). Cenozoic fossil mollusks from western Pacific islands; Gastropods (Eratoidae through Harpidae). U.S. Geological Survey Professional Paper. 533: i–iv, 1–84, pls 1–23 This article incorporates text from this source, which is in the public domain.
Ponder W.F. (1983). A revision of the Recent Xenophoridae of the World and of the Australian Fossil Species (Mollusca : Gastropoda). Memoir 17. The Australian Museum Sydney, Australia

[Fischer1807-1] Fischer von Waldheim, Gotthelf (1807). Muséum Demidoff, ou catalogue systématique et raisonné des curiosités de la nature et de l'art. Vol. 3: Végétaux et animaux. p. 213.

[Worms-2] WoRMS (2012). Xenophora. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=138664 on 2012-06-28

[3] Xenophora.org website [1] accessed 24 April 2014

[Kreipl_&_Alf-4] Kreipl, K. & Alf, A. (1999): Recent Xenophoridae. 148 pp. incl. 28 color plts. ConchBooks, Hackenheim, ISBN 3-925919-26-0.

[5] Powell A. W. B. (1979). New Zealand Mollusca, William Collins Publishers Ltd, Auckland, New Zealand. ISBN 0-00-216906-1

[6] OBIS Indo-Pacific Molluscan Database

[FW-7] Paleobiology Database

[8] Stromboidea

[9] G. Manganelli, V. Spadini, S. Cianfanelli The xenophorid gastropods of the Mediterranean Pliocene: the record of the Siena Basin^[usurped]

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Xenophora Temporal range: Late Cretaceous–Recent PreꞒ Ꞓ O S D C P T J K Pg N

A ventral view of a shell of Xenophora pallidula, with attached gastropod shells

A lateral view of the same shell
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Caenogastropoda
Order:	Littorinimorpha
Family:	Xenophoridae
Genus:	Xenophora Fischer von Waldheim, 1807^[1]
Species
See text
Synonyms
Phorus Montfort, 1810 Xenophora (Austrophora) Kreipl, Alf & Kronenberg, 1999· accepted, alternate representation † Xenophora (Endoptygma) Gabb, 1877 · accepted, alternate representation Xenophora (Xenophora) Fischer von Waldheim, 1807· accepted, alternate representation Xenophorus (incorrect subsequent spelling of Xenophora Fischer von Waldheim, 1807)